Elodium

(Sullivant) Austin

Musci Appalach., 52. 1870.

Etymology: Greek helos, marsh or marsh-meadow, and Latin -ium, diminutive, alluding to habitat
Basionym: Hypnum sect. Elodium Sullivant in A. Gray, Manual ed. 2, 668. 1856 Helodium name conserved
Treatment appears in FNA Volume 28. Treatment on page 321. Mentioned on page 323, 648.
 TaxonIllustrator 
HeloElodiumBlandowii.jpegElodium blandowiiPatricia M. Eckel

Plants medium-sized to large, green to yellowbrown. Stems erect or spreading, regularly ± 1-pinnate; paraphyllia filiform and narrowly lanceolate-foliose, branched or sometimes simple, cells long-rectangular, to 20: 1, smooth, apical cell-walls thin or somewhat thicker at tip; pseudoparaphyllia lanceolate-foliose, margins serrulate, proximal cells subquadrate, short to long-rectangular, smooth. Stem-leaves erect or erect-spreading, ovatelanceolate to rounded-cordate; margins irregularly revolute proximally or strongly, broadly or narrowly revolute to apex, entire and minutely toothed near apex, or denticulate throughout; apex abruptly to gradually narrowed, acumen broad or slender; costa surface smooth or with 1 or 2 teeth near insertion; laminal cells with abaxial surface roughened, 1-papillose centrally over lumen or more often near distal ends, papillae small, delicate; medial cells elongate; apical cell 6–7: 1, variously somewhat thickened distally, sometimes with one low papilla. Branch leaves similar. Sexual condition autoicous; perichaetia inner leaves cilia absent. Seta 2–6 cm. Capsule with stomata present in neck. Spores (10–) 11–15 (–16) µm.

Distribution

North America, Eurasia

Discussion

Species 3 (3 in the flora).

Elodium is distinguished from species of Thuidiaceae by its 1-pinnate branches with narrowly tapering branch ends. The marginal laminal cells in the often twisted acumen of species of Elodium are shorter than the cells mid acumen and produce a smooth, short apical cell. The basal laminal cells are larger, laxer, and paler than the more distal cells; in Echinophyllum they are thick-walled and short-rectangular to (sub)quadrate and concolorous. The perichaetial leaves of Elodium are nearly smooth throughout and bordered by minute, uniform denticulations, or sharply, distantly and irregularly toothed from distal cell ends. In Echinophyllum, the margins are sparsely short-ciliate with hyaline, long-celled, narrow teeth composed of (1 or) 2 elongate cells, free but parallel to the edge of the lamina. Both Elodium elodioides and E. paludosum seem intermediate in habitat preferences and able to tolerate seasonal desiccation.

Selected References

None.

Key

1 Stems with central strand absent; stems closely, evenly pinnate, branches nearly equal; leaf apices short-acuminate; setae 4-6 cm; capsules 3-4 mm; paraphyllia to 27 cells in length. Elodium blandowii
1 Stems with central strand present; stems loosely subpinnate, branches unequal; leaf apices long-acuminate; setae 2-2.8 cm; capsules 2-2.5 mm; paraphyllia 7-11 cells in length > 2
2 Leaves broadly cordate-ovate; apices abruptly acuminate; laminal cell walls thick, especially in margins, abaxial surface strongly 1-papillose centrally or nearer distal ends of lumen; leaf costa proximal abaxial surfaces with 1 or 2 scattered teeth. Elodium elodioides
2 Leaves ovate-lanceolate; apices gradually acuminate; laminal cell walls thin, especially in margins, abaxial surface smooth or weakly papillose-prorulose; leaf costa proximal abaxial surfaces smooth. Elodium paludosum
... more about "Elodium"
slender +
narrowed +
thicker +  and thin +
autoicous +, dioicous +, sexual +  and asexual +
paraphyllialike +
Patricia M. Eckel +
(Sullivant) Austin +
Hypnum sect. Elodium +  and Helodium +
autoicous +  and sexual +
cucullate +
strongly inclined;horizontal +
cylindric +
long-rectangular +
differentiated +
appendiculate +
nodulose +
slender +
North America +  and Eurasia +
Greek helos, marsh or marsh-meadow, and Latin -ium, diminutive, alluding to habitat +
oblong-lanceolate +
entire +  and serrulate +
denticulate +  and toothed +
apiculate +, blunt +, rostellate-rostrate +  and conic +
simple +, branched +  and lanceolate-foliose +
Musci Appalach., +
papillose +  and smooth +
lanceolate +
reddish orange +
elongate +
2 cm20 mm <br />0.02 m <br /> (6 cm60 mm <br />0.06 m <br />) +
globose +
plicate-sulcate +
erect-spreading;erect +
ovatelanceolate +  and rounded-cordate +
not stipitate +
spreading +  and erect +
Elodium +
Helodiaceae +
incrassate +
green +  and yellowbrown +
medium-sized;large +