Epilobium canum

(Greene) P. H. Raven

Ann. Missouri Bot. Gard. 63: 335. 1977.

Basionym: Zauschneria cana Greene Pittonia 1: 28. 1887
Treatment appears in FNA Volume 10.

Herbs suffruticose or not, with basal shoots from ± woody caudex, often decussate scales at base. Stems erect to ascending, often clumped but not matted, green or gray-green, terete, 10–110 (–120) cm, usually well-branched throughout, sometimes simple, strigillose and/or long-villous, usually mixed glandular puber­ulent distally, rarely glabrate. Leaves ± densely spaced, alternate and often fasciculate distally, subsessile, blade grayish green or green to silvery-canescent, usually narrowly linear to lanceolate or elliptic to ovate, rarely orbiculate, 0.6–5 (–6) × 0.1–2.5 cm, base cuneate to attenuate, margins subentire to sharply toothed, 3–15 teeth per side, veins inconspicuous or prominent, 3–7 per side, apex acute, sometimes with caducous dark mucro, surfaces usually ± densely strigillose, some­times mixed villous and/or glandular puberulent, rarely gla­brate; bracts much smaller and narrower. Inflorescences erect spikes or racemes, loose to congested, often branched, glandular puberulent and sometimes mixed strigillose or villous. Flowers: buds 11–18 × 4–6 mm, subsessile or pedicels 1–2 mm; floral-tube same color as petals, 16–32 × 5–8 mm, base slightly bulbous, ring of 8 irregular scales at base of filaments 4–6.5 mm from base inside; sepals same color as petals, 7–15 × 3.5–5 mm, abaxial surface densely pubescent; petals usually orange-red, very rarely white, obcordate, 8–17 × 5–9.5 mm, apical notch 2–3 mm; filaments light orange-red to white, those of longer stamens 12.5–32 mm, those of shorter ones 10–25 mm; anthers 2.7–4× 0.8–1.2 mm, apiculate; ovary 8–15 mm, glandular puberulent, often mixed villous; style light orange-red, 42–65 mm, glabrous, stigma 4-lobed, 1–1.4 × 2.4–3 mm, exserted 8–15 mm beyond anthers. Capsules straight or ± curved-ascending, 15–35 mm, sometimes beaked, surfaces glandular puberulent and strigillose; subsessile or pedicel 0–3 mm. Seeds broadly to narrowly obo­void, with constriction 0.6–0.8 mm from micropylar end, 1.5–2.6 × 0.9–1.3 mm, chalazal collar inconspicuous, light-brown, surface low papillose; coma easily detached, dingy white, 5.5–7 mm.

Distribution

sw United States, nw Mexico

Discussion

Subspecies 3 (3 in the flora).

This treatment recognizes three self-compatible but highly outcrossing subspecies marked by distinct but sometimes intergrading morphology and overlapping geographical ranges. R. N. Bowman and P. C. Hoch (1979) agreed with the treatment of Epilobium canum subsp. garrettii (n = 15) and subsp. latifolium (n = 30) by P. H. Raven (1976), but considering the complex intergrading patterns of variation involving the rest of this species, they combined the two remaining tetraploid subspecies recognized by Raven (subspp. angustifolium and mexicanum) with the remaining diploid subspecies into a single polyploid subsp. canum.

Selected References

None.

Key

1 Leaf blades linear to narrowly lanceolate or elliptic, rarely to narrowly ovate, 0.6–4.5 × 0.1–0.6(–0.8) cm, usually less than 0.6 cm wide, often fascicled distally, herbaceous to suffrutescent; stems 20–110(–120) cm. Epilobium canum subsp. canum
1 Leaf blades lanceolate to broadly ovate, 0.8–5(–6) × 0.4–2.3 cm, usually more than 0.6 cm wide, not fascicled; herbaceous; stems 10–50(–70) cm. > 2
2 Leaf blades usually lanceolate to ovate or broadly elliptical, rarely orbiculate, not coriaceous, margins subentire to distinctly denticulate, lateral veins obscure to conspicuous. Epilobium canum subsp. latifolium
2 Leaf blades ovate to broadly elliptical, coriaceous, margins prominently denticulate, lateral veins conspicuous. Epilobium canum subsp. garrettii

"narrower" is not a number.

... more about "Epilobium canum"
0.27 cm2.7 mm <br />0.0027 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
episepalous +
subequal +  and longer +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Peter C. Hoch +
(Greene) P. H. Raven +
4mm +  and 6.5mm +
cuneate +  and attenuate +
Zauschneria cana +
indehiscent +  and loculicidal +
5 cm50 mm <br />0.05 m <br /> (6 cm60 mm <br />0.06 m <br />) +
green +  and grayish green +
0.6 cm6 mm <br />0.006 m <br /> (5 cm50 mm <br />0.05 m <br />) +
toothed +  and entire +
silvery-canescent +
orbiculate +, usually narrowly linear +  and lanceolate or elliptic +
0.1 cm1 mm <br />0.001 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
smaller +
1.1 cm11 mm <br />0.011 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
recurved +  and erect +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
sessile +  and pedicellate +
curved-ascending +
subclavate +  and fusiform +
1.5 cm15 mm <br />0.015 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
subsessile +  and short-pedicellate +
dingy white +
persistent +
detached +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
sw United States +  and nw Mexico +
light orange-red +  and white +
basifixed +  and versatile +
1 cm10 mm <br />0.01 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
5mm +  and 8mm +
suberect +  and spread +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
perennial +
not +  and suffruticose +
branched +, loose +  and congested +
villous +, strigillose +, mixed +  and puberulent +
alternate +  and spaced +
deciduous +
decurrent +
subentire +  and sharply toothed +
villous +  and puberulent +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
numerous +  and 1 +
multicellular +
0 cm0 mm <br />0 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
white +  and orange-red +
8mm +  and 17mm +
obcordate +
0.5 cm5 mm <br />0.005 m <br /> (0.95 cm9.5 mm <br />0.0095 m <br />) +
Ann. Missouri Bot. Gard. +
branched +, loose +  and congested +
villous +, strigillose +, mixed +  and puberulent +
inflated +
basal +  and cauline +
clavate +  and globose +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
sculptured +  and smooth +
obovoid +
constricted +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
lanceolate +, , +, flushed with red or flushed with cream +  and green +
deciduous +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
branched +, loose +  and congested +
villous +, strigillose +, mixed +  and puberulent +
2 times as many as sepals +
simple +  and well-branched +
mixed +  and clumped +
110 cm1,100 mm <br />1.1 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
gray-green +  and green +
prostrate +  and decumbent +
erect +  and ascending +
glabrate +, long-villous +  and strigillose +
exfoliating +
10 cm100 mm <br />0.1 m <br /> (110 cm1,100 mm <br />1.1 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.24 cm2.4 mm <br />0.0024 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
light orange-red +
4.2 cm42 mm <br />0.042 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
aquatic +, amphibious +  and terrestrial +
strigillose +, puberulent +, glandular +, villous +  and mixed +
papillose +
constricted +
Epilobioideae alph. +, Boisduvaliinae raimann +  and Epilobiinae torrey +
Epilobium canum +
Epilobium sect. Zauschneria +
species +
3(-5)-aperturate +
prominent +  and inconspicuous +