Epilobium glaberrimum

Barbey in W. H. Brewer et al.

Bot. California 1: 220. 1876.

Common names: Glaucous willowherb
Synonyms: Epilobium fastigiatum var. glaberrimum (Barbey) Piper
Treatment appears in FNA Volume 10.

Herbs with branching, wiry basal shoots with crowded, often dark, scales. Stems many, erect or ascending, often densely clumped, slender, terete, 5–75 (–85) cm, simple or dis­tally branched, glabrous and glaucous, sometimes strigillose and/or mixed glandular puberulent distally, rarely with faint raised strigillose lines decurrent from margins of distal petioles. Leaves opposite and crowded proximal to inflorescence, alternate and widely spaced distally, subsessile and ± clasping; blade light green or blue-green, very narrowly lanceolate to narrowly ovate or elliptic, (0.5–) 1–7.2 × (0.3–) 0.5–1.8 cm, base cuneate, margins subentire or faintly denticulate, 8–15 teeth per side, veins inconspicuous, 3–6 per side, apex obtuse proximally to acute distally, glabrous and glaucous; bracts much reduced and narrower. Inflorescences erect racemes, glabrous or ± sparsely glandular puberulent, sometimes mixed strigillose. Flowers erect; buds 2.5–5 × 2–3.5 mm; pedicel 2–11 (–20) mm; floral-tube 0.7–2.3 × 1–2.5 mm, ring of spreading hairs at mouth inside or without a ring; sepals often red-tipped, sometimes keeled, 2.5–7.5 × 0.8–2 mm, abaxial surface glabrous or strigillose and/or glandular puberulent; petals usually light pink to rose-purple, rarely white, 3.4–10 (–12) × 1.8–5 mm, apical notch 0.5–3 mm; filaments purple to white or cream, those of longer stamens 2.4–5 (–6) mm, those of shorter ones 1.4–4 mm; anthers light yellow, 0.5–1.2 (–1.5) × 0.3–0.7 mm; ovary 10–42 mm, subglabrous to sparsely mixed strigillose and glandular puberulent; style light yellow or cream, 1.7–5 (–7) mm, stigma clavate to subcapitate, 0.8–3 × 0.5–1.2 mm, usually surrounded by, rarely exserted beyond, longer anthers. Capsules 15–75 mm, with small protuberance at base, surfaces subglabrous to sparsely mixed strigillose and glandular puberulent; pedicel 5–18 (–25) mm. Seeds narrowly obovoid, 0.7–1 (–1.3) × 0.3–0.5 mm, chalazal collar inconspicuous, light-brown or gray, surface coarsely papillose in longitudinal rows; coma readily detached, white, 4–9 mm.

Distribution

w North America, nw Mexico

Discussion

Subspecies 2 (2 in the flora).

Epilobium glaberrimum is restricted mainly to the Sierra Nevada-Cascade region, but also (subsp. fastigiatum) in the northern Rocky Mountains. It is one of the more easily identified species due to its consistently glabrous and glaucous aspect. Although the distributions and elevational ranges of the two subspecies appear to overlap extensively, they are largely allopatric with subsp. glaberrimum occurring in lower to mid montane zones and subsp. fastigiatum in upper montane to subalpine zones in the southern part of its range, and in lower zones farther north.

Selected References

None.

Key

1 Stems 20–75(–85) cm; leaf blades (2–)3–7.2 cm, very narrowly lanceolate to lanceolate or narrowly elliptic (length/width ratio more than 4); capsules 45–75 mm; petals (4–)5–10(–12) mm. Epilobium glaberrimum subsp. glaberrimum
1 Stems 5–35 cm; leaf blades (0.5–)1–3.4 cm, lanceolate to narrowly ovate or elliptic (length/width ratio less than 4); capsules 15–45(–55) mm; petals 3.4–8 mm. Epilobium glaberrimum subsp. fastigiatum

"narrower" is not a number.

... more about "Epilobium glaberrimum"
puberulent +, strigillose +  and glabrous +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
light yellow +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
episepalous +
subequal +  and longer +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
glaucous +  and glabrous +
acute +  and obtuse +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Peter C. Hoch +
Barbey in W. H. Brewer et al. +
wiry +  and branching +
cuneate +
indehiscent +  and loculicidal +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
blue-green +  and light green +
1 cm10 mm <br />0.01 m <br /> (7.2 cm72 mm <br />0.072 m <br />) +
toothed +  and entire +
very narrowly lanceolate +  and narrowly ovate or elliptic +
0.5 cm5 mm <br />0.005 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
reduced +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
recurved +  and erect +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
sessile +  and pedicellate +
curved +  and straight +
narrowly subcylindric +  and narrowly clavate +
1.5 cm15 mm <br />0.015 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
sessile +  and pedicellate +
gray +  and light-brown +
persistent +
detached +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Glaucous willowherb +
w North America +  and nw Mexico +
basifixed +  and versatile +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
1mm +  and 2.5mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
perennial +  and annual +
suffrutescent +
strigillose +, puberulent +, glandular +  and glabrous +
spaced +, alternate +, crowded +  and opposite +
deciduous +
raised +  and faint +
strigillose +
decurrent +
denticulate +  and subentire +
puberulent +, subglabrous +  and sparsely mixed strigillose +
1 cm10 mm <br />0.01 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
numerous +  and 1 +
multicellular +
1.8 cm18 mm <br />0.018 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
white +, usually light pink +  and rose-purple +
3.4mm +  and 10mm +
obovate +  and obcordate +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Bot. California +
inflated +
basal +  and cauline +
clavate +  and globose +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
sculptured +  and smooth +
obovoid +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
deciduous +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
purple +  and white or cream +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
2 times as many as sepals +
branched +, dis +  and simple +
75 cm750 mm <br />0.75 m <br /> (85 cm850 mm <br />0.85 m <br />) +
prostrate +  and decumbent +
ascending +  and erect +
puberulent +, glandular +, mixed +, strigillose +, glaucous +  and glabrous +
exfoliating +
slender +
5 cm50 mm <br />0.05 m <br /> (75 cm750 mm <br />0.75 m <br />) +
not woody +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
clavate +  and subcapitate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
cream +  and light yellow +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
aquatic +, amphibious +  and terrestrial +
puberulent +, subglabrous +  and sparsely mixed strigillose +
papillose +
constricted +
Epilobium fastigiatum var. glaberrimum +
Epilobium glaberrimum +
Epilobium sect. Epilobium +
species +
3(-5)-aperturate +
inconspicuous +