Epilobium hallianum

Haussknecht

Monogr. Epilobium, 261. 1884. (as halleanum)

Common names: Glandular willowherb
Synonyms: Epilobium brevistylum var. pringleanum (Haussknecht) Jepson E. brevistylum var. subfalcatum (Trelease) Munz E. brevistylum var. tenue (Trelease) Jepson E. brevistylum var. ursinum (Parish ex Trelease) Jepson E. delicatum var. tenue Trelease E. glandulosum var. tenue (Trelease) C. L. Hitchcock E. pringleanum Haussknecht E. pringleanum var. tenue (Trelease) Munz E. ursinum Parish ex Trelease E. ursinum var. subfalcatum Trelease
Treatment appears in FNA Volume 10.

Herbs with small, 3–6 mm, round or oblong, compact turions 1–5 cm below-ground. Stems strict, erect, terete, 2–50 (–60) cm, rarely branched only in larger plants, subglabrous proximal to inflorescence except for raised strigillose lines decurrent from margins of petioles, or sometimes ± densely long-villous through­out with inconspicuous decurrent lines. Leaves opposite proximally, alternate on inflorescence, subsessile or proximally with petioles 1–1.5 mm; blade ovate proximally to lanceolate or narrowly elliptic distally, 0.5–4.7 × 0.2–1.4 cm, base rounded to cuneate, mar­gins subentire proximally to denticulate distally, 8–20 teeth per side, veins inconspicuous, 3–6 per side, apex obtuse to subacute, surfaces mostly glabrous with strigillose margins; bracts much reduced. Inflorescences usually nodding in bud, erect later, open racemes, sometimes congested, usually mixed strigillose and glandular puberulent, rarely also mixed villous, or rarely subglabrous. Flowers erect; buds 2–5 × 1–2 mm; pedicel 3–8 mm; floral-tube 0.5–1.7 × 0.8–1.6 mm, with slightly raised ring of spreading hairs at mouth inside; sepals green, 1.2–2.8 × 0.5–1 mm, abaxial surface subglabrous or sparsely glandular puberulent; petals white, often fading pink, 1.6–5.5 × 1.2–3 mm, apical notch 0.3–1.2 mm; filaments white or cream, those of longer stamens 0.6–2.5 mm, those of shorter ones 0.4–1.5 mm; anthers cream, 0.2–0.9 × 0.2–0.5 mm; ovary 10–14 mm, strigillose and glandular puberulent or subglabrous; style cream, 0.8–5 mm, stigma clavate, entire, 0.4–1.2 × 0.3–0.7 mm, usually surrounded by, rarely exserted beyond, anthers. Capsules very narrowly cylindrical, (15–) 24–60 mm, surfaces usually subglabrous to mixed strigillose and glandular puberulent, rarely sparsely villous; pedicel 8–40 mm. Seeds narrowly fusiform to narrowly obovoid, 1.1–1.6 × 0.4–0.6 mm, chalazal collar ± conspicuous, 0.05–0.2 mm, light-brown, surface papillose; coma easily detached, white, 3–6 mm. 2n = 36.


Phenology: Flowering Jun–Sep.
Habitat: Semi-shaded stream banks, wet grassy slopes or meadows, bogs, seasonally wet sites, vernal pools.
Elevation: 100–3700 m.

Distribution

Alta., B.C., Sask., Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., S.Dak., Utah, Wash., Wyo.

Discussion

Epilobium hallianum has condensed fleshy turions and generally strict habit, suggesting an affinity with E. ciliatum or E. saximontanum, both with the AA arrangement. However, E. hallianum has the BB arrangement (S. R. Seavey and P. H. Raven 1978) and apparently more distant relationship to those other species.

In different parts of its geographical range Epilobium hallianum shows considerable morpho­logical variability, especially in leaf shape and margins, and in type and pattern of stem vestiture, including plants in the southern part of its range with densely villous stems that have been treated as E. ursinum. In part because of this variability and in part because the very characteristic condensed turions are easily lost during collection and/or often overlooked, E. hallianum is frequently misidentified.

Selected References

None.

Lower Taxa

None.
... more about "Epilobium hallianum"
puberulent +, glandular +  and subglabrous +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
subequal +  and longer +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
obtuse +  and subacute +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Peter C. Hoch +
Haussknecht +
subentire +, rounded +  and cuneate +
indehiscent +  and loculicidal +
0.5 cm5 mm <br />0.005 m <br /> (4.7 cm47 mm <br />0.047 m <br />) +
toothed +  and entire +
elliptic +, lanceolate +  and ovate +
0.2 cm2 mm <br />0.002 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
recurved +  and erect +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
sessile +  and pedicellate +
1.5 cm15 mm <br />0.015 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
curved +  and straight +
cylindrical +
2.4 cm24 mm <br />0.024 m <br /> (6 cm60 mm <br />0.06 m <br />) +
sessile +  and pedicellate +
persistent +
detached +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Glandular willowherb +
Alta. +, B.C. +, Sask. +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, S.Dak. +, Utah +, Wash. +  and Wyo. +
100–3700 m. +
basifixed +  and versatile +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
0.8mm +  and 1.6mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Semi-shaded stream banks, wet grassy slopes or meadows, bogs, seasonally wet sites, vernal pools. +
perennial +  and annual +
suffrutescent +
below-ground +
erect +  and nodding +
subglabrous +, villous +, mixed +, puberulent +, strigillose +  and long-villous +
proximally +  and subsessile +
alternate +  and opposite +
deciduous +
inconspicuous +
strigillose +
decurrent +
strigillose +
subglabrous +, puberulent +, glandular +  and strigillose +
1 cm10 mm <br />0.01 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
numerous +  and 1 +
multicellular +
0.8 cm8 mm <br />0.008 m <br /> (4 cm40 mm <br />0.04 m <br />) +
fading pink +  and white +
1.6mm +  and 5.5mm +
obovate +  and obcordate +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Flowering Jun–Sep. +
Monogr. Epilobium, +
inflated +
basal +  and cauline +
clavate +  and globose +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
sculptured +  and smooth +
narrowly fusiform +  and narrowly obovoid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
deciduous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
cream +  and white +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
2 times as many as sepals +
50 cm500 mm <br />0.5 m <br /> (60 cm600 mm <br />0.6 m <br />) +
prostrate +  and decumbent +
subglabrous +
exfoliating +
2 cm20 mm <br />0.02 m <br /> (50 cm500 mm <br />0.5 m <br />) +
not woody +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
aquatic +, amphibious +  and terrestrial +
villous +, puberulent +, usually subglabrous +  and mixed strigillose +
papillose +
constricted +
Epilobium brevistylum var. pringleanum +, E. brevistylum var. subfalcatum +, E. brevistylum var. tenue +, E. brevistylum var. ursinum +, E. delicatum var. tenue +, E. glandulosum var. tenue +, E. pringleanum +, E. pringleanum var. tenue +, E. ursinum +  and E. ursinum var. subfalcatum +
Epilobium hallianum +
Epilobium sect. Epilobium +
species +
3(-5)-aperturate +
inconspicuous +
denticulate +
oblong +  and round +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
inconspicuous +
denticulate +