familyOnagraceae
subfamilyOnagraceae subfam. Onagroideae
tribeOnagraceae tribe Epilobieae
genusEpilobium
sectionEpilobium sect. Epilobium
speciesEpilobium hornemannii
subspeciesEpilobium hornemannii subsp. behringianum

Epilobium hornemannii subsp. behringianum

(Haussknecht) Hoch & P. H. Raven

Ann. Missouri Bot. Gard. 64: 136. 1977.

Peter C. Hoch
Basionym: Epilobium behringianum Haussknecht Monogr. Epilobium, 277. 1884
Synonyms: E. alpinum var. behringianum (Haussknecht) S. L. Welsh E. bongardii Haussknecht
Treatment appears in FNA Volume 10.

Leaves ± coriaceous, petiole 4–9 mm proximally; cauline blade 1.5–6.2 × 0.7–2.2 cm. Flowers: sepals bright red, 5–7 × 2–2.5 mm; petals magenta to light pink, 8–10 (–11) × 3–6 mm. Capsules 35–55 mm. Seeds 0.9–1.6 × 0.4–0.5 mm, surfaces reticulate (smooth). 2n = 36.


Phenology: Flowering Jun–Aug.
Habitat: Moist meadows, seeps, stream banks, mossy or rocky crevices, usually coastal.
Elevation: 0–500 m.

Distribution

B.C., Yukon, Alaska, Asia (Russian Far East)

Discussion

Subspecies behringianum is a very distinctive coastal variant of the species, essentially replacing it in mari­time areas from northern British Columbia, including Haida Gwaii (the Queen Charlotte Islands), through Pacific coastal Alaska, and in similar habitats along the northwestern Pacific coast in Russia.

Selected References

None.

Lower Taxa

None.
... more about "Epilobium hornemannii subsp. behringianum"
RDF feed
2n chromosome quantity
36 +
Abaxial surface architecture or function or pubescence
glandular +
Abaxial surface pubescence
puberulent +  and strigillose +
Androecium position
epigynous +
Anther coloration
light yellow +
Anther fixation
versatile +
Anther length
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Anther orientation
erect +
Anther position
exserted +
Anther size
subequal +  and longer +
Anther width
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
Apex shape
obtuse +  and subacute +
Apical notch some measurement
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
Author
Peter C. Hoch +
Authority
(Haussknecht) Hoch & P. H. Raven +
Base architecture
bulbous +
Base life cycle
anthesis +
Base shape
attenuate +  and cuneate or rounded +
Basionyms
Epilobium behringianum +
Berry architecture or shape
nutlike +
Berry dehiscence
indehiscent +  and loculicidal +
Blade length
1.5 cm15 mm <br />0.015 m <br /> (6.2 cm62 mm <br />0.062 m <br />) +
Blade margin shape
toothed +  and entire +
Blade shape
broadly elliptic +  and spatulate proximally +
Blade texture
not +  and coriaceous +
Blade width
0.7 cm7 mm <br />0.007 m <br /> (2.9 cm29 mm <br />0.029 m <br />) +
Bract size
reduced +
Bracteole presence
absent +
Bud length
0.2 cm2 mm <br />0.002 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
Bud orientation
recurved +  and erect +
Bud width
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Bulblet growth form
cespitose +
Capsule architecture
sessile +  and pedicellate +
Capsule architecture or dehiscence
splitting +
Capsule architecture or shape
nutlike +
Capsule course
curved +  and straight +
Capsule dehiscence
dehiscent +
Capsule shape
narrowly subcylindric +  and narrowly clavate +
Capsule some measurement
3.5 cm35 mm <br />0.035 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
Carpel quantity
Many (?) +
Cauline blade length
1.5 cm15 mm <br />0.015 m <br /> (6.2 cm62 mm <br />0.062 m <br />) +
Cauline blade width
0.7 cm7 mm <br />0.007 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
Cel reproduction
vegetative +
Central column architecture
sessile +  and pedicellate +
Central column condition
intact +
Central column dehiscence
disintegrating +
Central column duration
persistent +
Chalazal collar coloration
brown +
Chalazal collar height or length or size
short +
Chalazal collar some measurement
0.05mm +  and 0.1mm +
Coma coloration
dingy white +
Coma duration
persistent +
Coma fusion
detached +
Coma presence
absent +
Coma some measurement
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Distribution
B.C. +, Yukon +, Alaska +  and Asia (Russian Far East) +
Elevation
0–500 m. +
Embryo-sac coating
oily +
Embryo-sac course
straight +
Embryo-sac presence
absent +
Embryo coating
oily +
Embryo course
straight +
Embryo presence
absent +
Endosperm coating
oily +
Endosperm course
straight +
Endosperm presence
absent +
Filament architecture
dithecal +
Filament fixation
basifixed +  and versatile +
Filament fusion
distinct +
Floral-tube architecture
not bulbous +
Floral-tube duration
deciduous +
Floral-tube length
0.1 cm1 mm <br />0.001 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
Floral-tube presence
absent +
Floral-tube quantity
, +
Floral-tube width
1.3mm +  and 2.8mm +
Flower architecture
actinomorphic +
Flower architecture or arrangement or growth form
solitary +
Flower life cycle
flowering +
Flower orientation
erect +
Flower part duration
deciduous +
Flower part life cycle
anthesis +
Flower reproduction
bisexual +
Fruit architecture
sessile +  and pedicellate +
Fruit architecture or dehiscence
splitting +
Fruit course
curved +  and straight +
Fruit dehiscence
dehiscent +
Fruit shape
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
Fruit size
slender +
Habitat
Moist meadows, seeps, stream banks, mossy or rocky crevices, usually coastal. +
Herb architecture
caulescent +
Herb duration
perennial +  and annual +
Herb growth form
suffrutescent +
Inflorescence orientation
nodding +  and erect +
Inflorescence position
axillary +
Inflorescence position or structure subtype
terminal +
Leaf architecture
simple +
Leaf architecture or arrangement
fasciculate +
Leaf arrangement
alternate +  and opposite +
Leaf duration
deciduous +
Leaf position
proximal +
Leaf texture
coriaceous +
Line prominence
raised +
Line pubescence
strigillose +
Line shape
decurrent +
Lobe quantity
Many (?) +
Locule quantity
1 +
Margin shape
denticulate +  and subentire +
Nectary presence
absent +
Nectary prominence
raised +
Number of lower taxa
0 +
Ovary architecture or function or pubescence
glandular +
Ovary architecture or structure in adjective form
4-locular +
Ovary arrangement
mixed +
Ovary position
inferior +
Ovary pubescence
strigillose +  and puberulent +
Ovary some measurement
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Ovule architecture
bitegmic +
Ovule arrangement or growth form
clustered +
Ovule orientation
anatropous +
Ovule quantity
numerous +  and 1 +
Papilla architecture
multicellular +
Pedicel atypical some measurement
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
Pedicel some measurement
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Perianth position
epigynous +
Petal architecture or shape
emarginate +
Petal arrangement
imbricate +
Petal arrangement or shape
convolute +
Petal atypical length
1 cm10 mm <br />0.01 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Petal coloration
magenta +  and light pink +
Petal length
8mm +  and 10mm +
Petal presence
absent +
Petal quantity
4 +
Petal shape
obovate +  and obcordate +
Petal width
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Petiole architecture
subsessile +
Petiole some measurement
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Phenology
Flowering Jun–Aug. +
Pollen-grain dehiscence or orientation
longitudinal +
Pollen fixation
basifixed +
Polyad arrangement
singly +
Publication title
Ann. Missouri Bot. Gard. +
Publication year
1977 +
Raceme architecture
open +
Raceme architecture or function or pubescence
glandular +
Raceme arrangement
mixed +
Raceme pubescence
puberulent +  and strigillose +
Raphide quantity
abundant +
Reference
None +
Rim shape
inflated +
Ring count or density
sparse +
Ring presence
absent +
Ring prominence
raised +
Rosette position
basal +  and cauline +
Row quantity
1 +
Secretion condition
wet +
Secretion condition or texture
dry +
Secretion relief
papillate +
Secretion shape
clavate +  and globose +
Seed length
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
Seed quantity
many +
Seed relief
sculptured +  and smooth +
Seed shape
oblanceoloid +  and fusiform +
Seed width
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
Sepal arrangement or dehiscence
valvate +
Sepal coloration
bright red +
Sepal duration
deciduous +
Sepal length
? (?) +
Sepal orientation
spreading +
Sepal quantity
4 +
Sepal width
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Septum architecture
parenchymatous +
Septum arrangement
parietal +, axile +  and placentation +
Septum presence
absent +
Septum width
thin +
Series length or size
longer +
Series quantity
2 +
Series size
unequal +
Series variability
equal +
Set length or size
longer +
Set quantity
2 +
Set variability
equal +
Shrub architecture
unarmed +
Shrub growth form
not clonal +
Shrub habitat
aquatic +, amphibious +  and terrestrial +
Slit dehiscence or orientation
longitudinal +
Sobole architecture
leafy +
Sobole architecture or pubescence
scaly +
Sobole height or length or size
short +
Sobole location
epigeous +
Species position
proximal +
Spike architecture
leafy +
Stamen arrangement
singly +
Stamen filament atypical some measurement
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Stamen filament coloration
cream +  and light pink +
Stamen filament some measurement
0.12 cm1.2 mm <br />0.0012 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Stamen quantity
8 +
Stamen size or quantity
2 times as many as sepals +
Stem architecture
branched +  and simple +
Stem architecture or function or pubescence
glandular +
Stem arrangement
mixed +  and clumped +
Stem growth form or orientation
prostrate +  and decumbent +
Stem orientation
ascending +  and erect +
Stem position
proximal +
Stem pubescence
puberulent +, strigillose +  and subglabrous +
Stem relief
exfoliating +
Stem shape
terete +
Stem some measurement
10 cm100 mm <br />0.1 m <br /> (45 cm450 mm <br />0.45 m <br />) +
Stem texture
not woody +
Stigma architecture or shape
entire +
Stigma coloration
cream +
Stigma length
0.12 cm1.2 mm <br />0.0012 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Stigma quantity
1 +
Stigma shape
cylindrical +  and clavate +
Stigma structure in adjective form
commissural +
Stigma width
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Stipule architecture
subsessile +  and petiolate +
Stipule duration
caducous +
Stipule position
intrapetiolar +
Stipule presence
absent +
Stipule size
small +
Stolon architecture
tipped +
Style coloration
cream +  and white +
Style orientation
erect +
Style quantity
1 +
Style some measurement
0.2 cm2 mm <br />0.002 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Subshrub architecture
unarmed +
Subshrub growth form
not clonal +
Subshrub habitat
aquatic +, amphibious +  and terrestrial +
Surface architecture or coloration or relief
reticulate +
Surface architecture or function or pubescence
glandular +
Surface arrangement
mixed +
Surface condition or texture
dry +
Surface pubescence
strigillose +, puberulent +, , +  and glabrous +
Surface quantity
19 (18) +
Surface relief
smooth +  and papillose +
Surface shape
ridged +
Surface size
constricted +
Synonyms
E. alpinum var. behringianum +  and E. bongardii +
Taxon family
Onagraceae +
Taxon name
Epilobium hornemannii subsp. behringianum +
Taxon parent
Epilobium hornemannii +
Taxon rank
subspecies +
Tetrad arrangement
singly +
Tetrad dehiscence
3(-5)-aperturate +
Tooth quantity
10 (25) +
Turion arrangement
decussate +
Turion texture
fleshy +
Vein prominence
inconspicuous +
Vein quantity
4 (7) +
Viscin thread dehiscence or orientation
longitudinal +
Viscin thread fusion
united +
Volume
Volume 10 +
Whole organism growth form
suffrutescent +