Epilobium hornemannii subsp. hornemannii

Synonyms: Epilobium cupreum Lange E. foucaudianum H. Léveillé E. steckerianum Fernald
Treatment appears in FNA Volume 10.

Leaves not coriaceous, petiole 3–7 mm proximally; cauline blade 1.5–5.5 × 0.7–2.9 cm. Flowers: sepals sometimes red-tipped, 2–4.5 × 1–2.2 mm; petals usually rose-purple to light pink, rarely white, 3–9 × 2–5.5 mm. Capsules 40–65 mm. Seeds 0.9–1.2 × 0.3–0.5 mm, surfaces distinctly papillose. 2n = 36.


Phenology: Flowering (May–)Jun–Aug.
Habitat: Banks of montane to alpine streams and lakes, open tussock meadows, willow swales, gravelly ridges, stabilized scree slopes, road­side ditches.
Elevation: 50–3700 m.

Distribution

Greenland, Alta., B.C., N.B., Nfld. and Labr. (Nfld.), N.W.T., N.S., Nunavut, Ont., Que., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Maine, Mont., Nev., N.H., N.Mex., N.Y., Oreg., S.Dak., Utah, Wash., Wyo., n Eurasia

Discussion

Subspecies hornemannii is one of the more com­mon and widely distributed members of the Alpinae alliance and carries the CC chromosomal arrangement. It grows from near sea level in the north (Alaska, Northwest Territories) to high alpine (to 3700 m) in the southern Rockies and Sierra Nevada; it also occurs in northern Europe, subarctic and Russian Far East, and Japan. In Nunuvut, it is found only on Charlton Island in James Bay. It often grows sympatrically with E. ciliatum and Epilobium saximontanum (both AA arrangement); occasional hybrids are found, but they often have much reduced seed fertility, probably due to chromosomal irregularities.

Subspecies hornemannii occurs occasionally in spring pools, forming distinctive dense floating mats of ascending stems. Specimens growing in very moist habitats often have pale, thin epigeal stolons with scales, rather than typical leafy soboles.

Selected References

None.

Lower Taxa

None.
puberulent +  and strigillose +
light yellow +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
subequal +  and longer +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
obtuse +  and subacute +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
Peter C. Hoch +
Reichenbach +
attenuate +  and cuneate or rounded +
Epilobium nutans +
indehiscent +  and loculicidal +
1.5 cm15 mm <br />0.015 m <br /> (6.2 cm62 mm <br />0.062 m <br />) +
toothed +  and entire +
broadly elliptic +  and spatulate proximally +
not +  and coriaceous +
0.7 cm7 mm <br />0.007 m <br /> (2.9 cm29 mm <br />0.029 m <br />) +
reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
recurved +  and erect +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
sessile +  and pedicellate +
curved +  and straight +
narrowly subcylindric +  and narrowly clavate +
4 cm40 mm <br />0.04 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (2.9 cm29 mm <br />0.029 m <br />) +
sessile +  and pedicellate +
dingy white +
persistent +
detached +
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Greenland +, Alta. +, B.C. +, N.B. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, N.S. +, Nunavut +, Ont. +, Que. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Maine +, Mont. +, Nev. +, N.H. +, N.Mex. +, N.Y. +, Oreg. +, S.Dak. +, Utah +, Wash. +, Wyo. +  and n Eurasia +
50–3700 m. +
basifixed +  and versatile +
0.1 cm1 mm <br />0.001 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
1.3mm +  and 2.8mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Banks of montane to alpine streams and lakes, open tussock meadows, willow swales, gravelly ridges, stabilized scree slopes, roadside ditches. +
perennial +  and annual +
suffrutescent +
nodding +  and erect +
alternate +  and opposite +
deciduous +
not coriaceous +
strigillose +
decurrent +
denticulate +  and subentire +
strigillose +  and puberulent +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
numerous +  and 1 +
multicellular +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
white +, usually rose-purple +  and light pink +
3mm +  and 9mm +
obovate +  and obcordate +
0.2 cm2 mm <br />0.002 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
Flowering (May–)Jun–Aug. +
Iconogr. Bot. Pl. Crit. +
puberulent +  and strigillose +
inflated +
basal +  and cauline +
clavate +  and globose +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
sculptured +  and smooth +
oblanceoloid +  and fusiform +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
deciduous +
0.1 cm1 mm <br />0.001 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
cream +  and light pink +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
2 times as many as sepals +
branched +  and simple +
mixed +  and clumped +
prostrate +  and decumbent +
ascending +  and erect +
puberulent +, strigillose +  and subglabrous +
exfoliating +
10 cm100 mm <br />0.1 m <br /> (45 cm450 mm <br />0.45 m <br />) +
not woody +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
cylindrical +  and clavate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
cream +  and white +
0.2 cm2 mm <br />0.002 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
aquatic +, amphibious +  and terrestrial +
strigillose +, puberulent +, , +  and glabrous +
papillose +
constricted +
Epilobium cupreum +, E. foucaudianum +  and E. steckerianum +
Epilobium hornemannii subsp. hornemannii +
Epilobium hornemannii +
subspecies +
3(-5)-aperturate +
inconspicuous +