Epilobium lactiflorum

Haussknecht

Oesterr. Bot. Z. 29: 89. 1879.

Common names: Milkflower willowherb épilobeà fleurs blanches
Synonyms: Epilobium alpinum var. lactiflorum (Haussknecht) C. L. Hitchcock E. canadense H. Léveillé E. canadense var. albescens H. Léveillé E. hornemannii var. lactiflorum (Haussknecht) D. Löve
Treatment appears in FNA Volume 10.

Herbs with short, leafy epigeal soboles. Stems ascending to sub­erect, often clumped, terete, 15–50 cm, usually simple, rarely branched proximally, subglabrous proximal to inflorescence except for raised densely strigillose lines decurrent from margins of petioles, usually mixed strigillose and glandular puberulent distally. Leaves opposite proximal to inflorescence or just proximal 1/3, alternate distally, petiole 3–12 mm, ± winged; blade broadly spatulate to ovate proxi­mally, narrowly ovate to narrowly lanceolate distally, 2–5.5 × 0.8–2.4 cm, base attenuate to cuneate, margins subentire proximally to denticulate distally with 7–16 teeth per side, more marked distally, lateral-veins inconspicuous, 4–8 per side, apex obtuse proximally to subacute distally, surfaces glabrous except for strigil­lose margins; bracts reduced and narrower. Inflorescences nodding in bud, later erect, ± open racemes, mixed strigillose and glandular puberulent. Flowers suberect; buds 2–5 × 1.5–3.5 mm; pedicel 5–15 mm; floral-tube 1–2.2 × 1–3 mm, inner surface glabrous without ring; sepals often flushed purplish red, frequently keeled, (2–) 3–5.5 × 0.9–1.8 mm, abaxial surface sparsely glandular puberulent, sometimes mixed strigillose; petals white, rarely with red veins or flushed light pink, 3–8.5 × 1.6–4.5 mm, apical notch 0.7–1.4 mm; fila­ments white to cream, those of longer stamens 1.4–4 mm, those of shorter ones 1.1–3 mm; anthers light yellow, 0.4–0.9 × 0.3–0.6 mm; ovary 20–40 mm, glandular puber­ulent; style cream or white, 1.4–4.6 mm, stigma clavate or rarely subcapitate and indented apically, entire, 1.2–2.5 × 0.4–1.6 mm, surrounded by anthers. Capsules slender, sometimes flushed reddish green, ± ascending, 50–100 mm, surfaces sparsely glandular puberulent; pedicel 15–45 mm. Seeds narrowly obo­void, 1.1–1.7 × 0.4–0.6 mm, chalazal collar 0.05–0.1 mm, blond or light-brown, surface reticulate or some­times barely rugose; coma easily detached, white, 7–14 mm. 2n = 36.


Phenology: Flowering Jun–Sep.
Habitat: Montane stream banks, moist crevices and ledges, gravelly roadsides, burned-over woodlands, sandy moraines, subalpine forests, alpine meadows.
Elevation: 50–3800 m.

Distribution

Greenland, Alta., B.C., N.B., Nfld. and Labr., N.W.T., N.S., Ont., Que., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Maine, Mont., Nev., N.H., N.Mex., Oreg., Utah, Vt., Wash., Wyo., Eurasia

Discussion

Epilobium lactiflorum has a nearly circum-subarctic distribution in North America (including coastal Greenland) and Eurasia, extending south into alpine and cool montane habitats along mountain axes. This distribution is similar to that of E. anagallidifolium and E. hornemannii (all with CC chromosomal arrangement), and these species commonly grow in similar habitats as well.

Petal color can be variable in many Epilobium species, but E. lactiflorum (white flowers) differs quite consistently from E. hornemannii (rose-purple to light pink or rarely white) in that feature. Mature fruits and pedicels are also fairly longer in E. lactiflorum. Despite their morphological similarities and broadly overlapping ranges and habitats, E. lactiflorum and E. hornemannii subsp. hornemannii do not appear to hybridize with much frequency, although intermediates, with only moderately reduced seed fertility, might be difficult to detect.

Selected References

None.

Lower Taxa

None.

"narrower" is not a number.

... more about "Epilobium lactiflorum"
strigillose +  and puberulent +
light yellow +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
episepalous +
subequal +  and longer +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
subacute +  and obtuse +
pink +  and flushed light +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Peter C. Hoch +
Haussknecht +
attenuate +  and cuneate +
indehiscent +  and loculicidal +
2 cm20 mm <br />0.02 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
toothed +  and entire +
narrowly ovate +  and narrowly lanceolate +
0.8 cm8 mm <br />0.008 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
recurved +  and erect +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
sessile +  and pedicellate +
green +  and flushed reddish +
curved +  and straight +
narrowly subcylindric +  and narrowly clavate +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
sessile +  and pedicellate +
persistent +
detached +
0.7 cm7 mm <br />0.007 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Milkflower willowherb +  and épilobeà fleurs blanches +
Greenland +, Alta. +, B.C. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Ont. +, Que. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Maine +, Mont. +, Nev. +, N.H. +, N.Mex. +, Oreg. +, Utah +, Vt. +, Wash. +, Wyo. +  and Eurasia +
50–3800 m. +
basifixed +  and versatile +
0.1 cm1 mm <br />0.001 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
1mm +  and 3mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Montane stream banks, moist crevices and ledges, gravelly roadsides, burned-over woodlands, sandy moraines, subalpine forests, alpine meadows. +
perennial +  and annual +
suffrutescent +
erect +  and nodding +
deciduous +
strigillose +
decurrent +
inconspicuous +
subentire +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
numerous +  and 1 +
multicellular +
1.5 cm15 mm <br />0.015 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
white +  and cream +
obovate +  and obcordate +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Flowering Jun–Sep. +
Oesterr. Bot. Z. +
puberulent +  and strigillose +
inflated +
basal +  and cauline +
clavate +  and globose +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
sculptured +  and smooth +
fusiform +  and narrowly ellipsoid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
red +  and flushed purplish +
deciduous +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
aquatic +, amphibious +  and terrestrial +
2 times as many as sepals +
branched +  and simple +
mixed +  and clumped +
prostrate +  and decumbent +
erect +  and ascending +
puberulent +, strigillose +  and subglabrous +
exfoliating +
decurrent +  and terete +
15 cm150 mm <br />0.15 m <br /> (50 cm500 mm <br />0.5 m <br />) +
not woody +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
indented +, subcapitate +  and clavate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
white +  and cream +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.46 cm4.6 mm <br />0.0046 m <br />) +
aquatic +, amphibious +  and terrestrial +
puberulent +  and glabrous +
constricted +
Epilobium alpinum var. lactiflorum +, E. canadense +, E. canadense var. albescens +  and E. hornemannii var. lactiflorum +
Epilobium lactiflorum +
Epilobium sect. Epilobium +
species +
3(-5)-aperturate +
denticulate +