Epilobium saximontanum

Haussknecht

Oesterr. Bot. Z. 29: 119. 1879.

Common names: Rocky Mountain willowherb épilobe des Rocheuses
Synonyms: Epilobium adenocaulon subsp. rubescens (Rydberg) Hiitonen E. drummondii Haussknecht E. drummondii var. latiusculum Rydberg E. latiusculum (Rydberg) Rydberg E. ovatifolium Rydberg E. rubescens Rydberg E. scalare Fernald E. stramineum Rydberg
Treatment appears in FNA Volume 10.

Herbs usually with sessile, fleshy, underground turions, or sometimes thick, elongated shoots with dark, decussate scales. Stems erect, strict, terete, 4–55 cm, simple or well branched in age, subglabrous proximally to mixed strigillose and glandular puberulent distally, with raised strigillose lines decurrent from margins of petioles. Leaves opposite proximal to inflorescence, alternate and reduced distally, often ± appressed, usually subsessile, rarely petiole 1–3 mm, often clasping; blade obovate proximally to ovate, lanceolate, or narrowly elliptic distally, 1–5.5 (–6.5) × 0.4–2 (–2.4) cm, base rounded or obtuse, margins low denticulate, 9–30 teeth per side, veins ± conspicuous, 3–6 per side, apex subacute, surfaces subglabrous with strigillose margins; bracts much reduced. Inflorescences erect, sometimes nodding in bud, racemes, sometimes sparsely branched. Flowers erect; buds 2–3.5 × 1.8–2.5 mm; pedicel 0–1 mm; floral-tube 0.8–1.4 × 0.8–1.9 mm, ring of sparse spreading hairs at mouth inside; sepals sometimes flushed red, 1.2–3.5 × 0.6–1.4 mm, abaxial surface strigillose and sometimes mixed glandular puberulent; petals usually white, infrequently pink, 2.2–5 (–7) × 1.7–3.2 mm, apical notch 0.4–1.5 mm; filaments usually cream, rarely light pink, those of longer stamens 2–3.5 mm, those of shorter ones 1–2 mm; anthers cream to light yellow, 0.3–0.8 × 0.3–0.5 mm; ovary 9–30 mm, densely strigillose and glandular puberulent; style cream or yellow, 1.6–2.8 mm, stigma usually narrowly to broadly clavate, rarely subcapitate, 1–3 × 0.8–2 mm, surrounded by at least longer anthers. Capsules 30–55 (–70) mm, surfaces mixed strigillose and glan­dular puberulent; usually subsessile, rarely pedicel 1–5 mm, often appressed to stem. Seeds very narrowly obovoid, 1–1.6 (–1.8) × 0.4–0.6 mm, chalazal collar 0.1–0.2 mm, light-brown or gray, surface rugose to papillose; coma usually readily detached, white, 3–9 mm. 2n = 36.


Phenology: Flowering Jul–Sep.
Habitat: Montane semi-shaded stream banks, damp meadows, mossy seeps, wet slatey cliffs, disturbed or seasonally damp areas.
Elevation: 0–3700 m.

Distribution

Alta., B.C., Man., Nfld. and Labr. (Nfld.), Ont., Que., Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., S.Dak., Utah, Wyo.

Discussion

Epilobium saximontanum is morphologically similar to E. ciliatum (especially subsp. glandulosum) with which it also shares the AA chromosome arrangement. However, in addition to its fleshy compact turions, it very characteristically has notably appressed capsules, unlike most other species in the genus, and a notably strict habit.

The distribution of Epilobium saximontanum is unusual; it includes the Rocky Mountain region, only barely reaching the high southern Sierra Nevada, dis­junct to the Black Hills of South Dakota, and more widely in eastern Canada, from the shores of Hudson Bay to Newfoundland. Specimens are fairly uniform across this wide and rather discontinuous range, although locally they show some variability, pos­sibly due to hybridization with any of several species that may be sympatric with it. H. Lewis and D. M. Moore (1962) reported hybrids between E. saximontanum (cited as E. brevistylum) and E. ciliatum subsp. ciliatum (cited as E. adenocaulon) from Colorado, and her­barium specimens with E. saximontanum and apparent hybrids are not uncommon.

Selected References

None.

Lower Taxa

None.
... more about "Epilobium saximontanum"
puberulent +  and strigillose +
cream +  and light yellow +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
episepalous +
subequal +  and longer +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
subacute +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Peter C. Hoch +
Haussknecht +
obtuse +  and rounded +
indehiscent +  and loculicidal +
5.5 cm55 mm <br />0.055 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
2 cm20 mm <br />0.02 m <br /> (2.4 cm24 mm <br />0.024 m <br />) +
1 cm10 mm <br />0.01 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
toothed +  and entire +
elliptic +, lanceolate +, ovate +  and obovate +
0.4 cm4 mm <br />0.004 m <br /> (2 cm20 mm <br />0.02 m <br />) +
reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
recurved +  and erect +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
sessile +  and pedicellate +
5.5 cm55 mm <br />0.055 m <br /> (7 cm70 mm <br />0.07 m <br />) +
curved +  and straight +
narrowly subcylindric +  and narrowly clavate +
3 cm30 mm <br />0.03 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
sessile +  and pedicellate +
gray +  and light-brown +
persistent +
detached +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
Rocky Mountain willowherb +  and épilobe des Rocheuses +
Alta. +, B.C. +, Man. +, Nfld. and Labr. (Nfld.) +, Ont. +, Que. +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, S.Dak. +, Utah +  and Wyo. +
0–3700 m. +
basifixed +  and versatile +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
0.8mm +  and 1.9mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Montane semi-shaded stream banks, damp meadows, mossy seeps, wet slatey cliffs, disturbed or seasonally damp areas. +
perennial +  and annual +
suffrutescent +
nodding +  and erect +
alternate +  and opposite +
deciduous +
reduced +
strigillose +
decurrent +
strigillose +
denticulate +
puberulent +  and strigillose +
0.9 cm9 mm <br />0.009 m <br /> (3 cm30 mm <br />0.03 m <br />) +
numerous +  and 1 +
multicellular +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
pink +  and white +
2.2mm +  and 5mm +
obovate +  and obcordate +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Flowering Jul–Sep. +
Oesterr. Bot. Z. +
inflated +
basal +  and cauline +
clavate +  and globose +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
sculptured +  and smooth +
obovoid +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
flushed red +
deciduous +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
elongated +
aquatic +, amphibious +  and terrestrial +
light pink +  and cream +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
2 times as many as sepals +
branched +  and simple +
prostrate +  and decumbent +
puberulent +, strigillose +  and subglabrous +
exfoliating +
decurrent +  and terete +
4 cm40 mm <br />0.04 m <br /> (55 cm550 mm <br />0.55 m <br />) +
not woody +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
yellow +  and cream +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
aquatic +, amphibious +  and terrestrial +
puberulent +, strigillose +  and subglabrous +
rugose +  and papillose +
constricted +
Epilobium adenocaulon subsp. rubescens +, E. drummondii +, E. drummondii var. latiusculum +, E. latiusculum +, E. ovatifolium +, E. rubescens +, E. scalare +  and E. stramineum +
Epilobium saximontanum +
Epilobium sect. Epilobium +
species +
3(-5)-aperturate +
underground +
conspicuous +