Eriophyllum lanatum var. achilleoides

(de Candolle) Jepson

Man. Fl. Pl. Calif., 1118. 1925.

Basionym: Bahia achillioides de Candolle in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 657. 1836 (as achillaeoides)
Synonyms: Eriophyllum lanatum var. aphanactis J. T. Howell
Treatment appears in FNA Volume 21. Mentioned on page 357, 358, 359, 360.

Perennials (sometimes flowering first-year; ± taprooted). Proximal leaves usually mostly alternate (opposite in ne California); blades oblanceolate to ovate, 1–2 (–3) -pinnately lobed (lobes usually ± triangular), ultimate margins often toothed (teeth 1–4, 1–2 mm), often crispate, usually revolute, abaxial faces woolly, adaxial less so or glabrate. Heads usually 3–8 per array, sometimes borne singly (n California, s Oregon). Peduncles mostly 3–10 cm. Involucres usually 8–10 mm diam. Ray-florets usually 9–13, sometimes 0; laminae usually 6–9 (–16) mm. Cypselae 2–3 (–3.3) mm; pappi (0.3–) 0.5–0.9 (–1) mm. 2n = 16, 32.


Phenology: Flowering Apr–Jun.
Habitat: Dry sites, chaparral, forests
Elevation: 30–1300 m

Discussion

In northern California, a taxonomically perplexing series of intermediate populations connects vars. arachnoideum, achillioides, and grandiflorum. Polyploiids characterize the regions where the varieties meet. Putative arachnoideum-achillioides hybrid derivatives, compared to var. achillioides, have smaller, broader, thinner, and more shallowly lobed leaves, and shorter pappi. Putative achillioides-grandiflorum hybrid derivatives, compared to var. achillioides, have longer peduncles, larger heads (often borne singly), and longer ray laminae. Variety aphanactis (2n = 32), an Inner Coast Range local, rayless variant, is probably of achillioides-grandiflorum parentage. Varieties achillioides, arachnoideum, grandiflorum, and lanceolatum overlap in northwestern California and southwestern Oregon. L. Constance (1937, p. 89), referring to the Klamath area, observed “that it seems quite impossible to assign many specimens definitely to any one subspecific category.” Variety achillioides passes gradually through northern California and southern Oregon populations into var. leucophyllum west of the Cascade Mountains. Oregon specimens with ternately lobed leaves and larger heads often borne singly (Eriophyllum ternatum Greene) are part of the continuum.

L. Constance (1937, p. 87) had var. achillioides, although “uncommon in the Sierra Nevada, occurring as far south as Mariposa County.”

Selected References

None.

Lower Taxa

None.
not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
Dale E. Johnson +  and John S. Mooring +
(de Candolle) Jepson +
decurrent +
Bahia achillioides +
compound +  and simple +
lobed;oblanceolate;ovate +
1 cm10 mm <br />0.01 m <br /> (8 cm80 mm <br />0.08 m <br />) +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
yellow +  and white +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.33 cm3.3 mm <br />0.0033 m <br />) +
glabrous +  and hairy +
papillate +
4(-5)-angled +, linear-clavate +  and prismatic flattened or 3-angled +
fertile +  and bisexual +
fertile +  and bisexual +
Calif. +, Nev. +  and Oreg. +
30–1300 m +
woolly +  and hairy +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Dry sites, chaparral, forests +
compound-corymbiform +, corymbiform +, discoid +  and radiate +
disciform +  and discoid +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
campanulate;hemispheric +
0.9 cm9 mm <br />0.009 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
golden yellow +  and yellow +
6mm;20[mm +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
2mm;7]mm +
sessile +  and petiolate +
papillate +
deltate +
dentate to pinnatifid or palmatifid +  and entire +
2-carpellate +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
persistent +
of 1 +  and 12+ aristate +
coroniform +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Flowering Apr–Jun. +
scarious-margined +
connate +  and distinct +
weakly +  and flat +
glabrous +, glabrate +, woolly +  and tomentose +
5 +  and 13 +
Man. Fl. Pl. Calif., +
pitted +  and smooth +
convex;conic +
6 +  and 12 +
lacerate +, cuneate +  and lanceolate or lance-linear +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
exalbuminous +
Illustrated +
erect +  and decumbent +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Eriophyllum lanatum var. aphanactis +
Eriophyllum lanatum var. achilleoides +
Eriophyllum lanatum +
variety +
campanulate +
crispate +  and toothed +
10 cm100 mm <br />0.1 m <br /> (100 cm1,000 mm <br />1 m <br />) +