Eriophyllum lanatum var. integrifolium

(Hooker) Smiley

Univ. Calif. Publ. Bot. 9: 378. 1921.

Endemic
Basionym: Trichophyllum integrifolium Hooker Fl. Bor.-Amer. 1: 316. 1833
Synonyms: Eriophyllum lanatum var. cuneatum (Kellogg) Jepson Eriophyllum lanatum var. monoense (Rydberg) Jepson
Treatment appears in FNA Volume 21. Treatment on page 360. Mentioned on page 358, 359.

Shrubs (multistemmed; taprooted). Proximal leaves alternate; blades cuneate, oblanceolate, or obovate, margins entire or ± toothed to lobed (lobes 3–5), plane or slightly revolute, faces (persistently) woolly. Heads usually borne singly. Peduncles mostly 3–10 cm. Involucres 6–10 mm diam. Ray-florets (5–) 8 (–10); laminae 6–12 mm. Cypselae (2.3–) 3–4 (–5) mm; pappi (0.3–) 0.5–1 (–2) mm (scales equal or unequal). 2n = 16, 32, 48, 64.


Phenology: Flowering Jun–Aug.
Habitat: Dry or rocky sites, sagebrush, conifer forests alpine fell fields
Elevation: 1200–3500 m

Distribution

V21-902-distribution-map.gif

Calif., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

L. Constance (1937) sorted the regional races of this complex, some taxonomically recognized, into five geographic groups. P. A. Munz (1959) referred high elevation populations in California and adjacent states to var. monoense (Rydberg) Jepson. Variety integrifolium intergrades with var. grandiflorum. Constance and also L. Abrams and R. S. Ferris (1923–1960, vol. 4) applied var. cuneatum (Kellogg) Jepson to individuals of what appear to be vars. grandiflorum × integrifolium parentage as well as to variants of vars. achillioides, grandiflorum, and integrifolium. Variety integrifolium intergrades with var. lanatum in Oregon and Washington near the Columbia River. The intermediate populations that have been analyzed are polyploid (J. S. Mooring 2001).

Selected References

None.

Lower Taxa

None.
64 +, 48 +, 32 +  and 16 +
not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
Dale E. Johnson +  and John S. Mooring +
(Hooker) Smiley +
decurrent +
Trichophyllum integrifolium +
compound +  and simple +
obovate;oblanceolate;obovate;oblanceolate;cuneate +
1 cm10 mm <br />0.01 m <br /> (8 cm80 mm <br />0.08 m <br />) +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
yellow +  and white +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
glabrous +  and hairy +
papillate +
4(-5)-angled +, linear-clavate +  and prismatic flattened or 3-angled +
fertile +  and bisexual +
fertile +  and bisexual +
Calif. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
1200–3500 m +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Dry or rocky sites, sagebrush, conifer forests alpine fell fields +
compound-corymbiform +, corymbiform +, discoid +  and radiate +
disciform +  and discoid +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
campanulate;hemispheric +
golden yellow +  and yellow +
6mm;20[mm +
0.6 cm6 mm <br />0.006 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
2mm;7]mm +
sessile +  and petiolate +
papillate +
deltate +
dentate to pinnatifid or palmatifid +  and entire +
more or less toothed +  and lobed plane or slightly revolute +
2-carpellate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
persistent +
of 1 +  and 12+ aristate +
coroniform +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Flowering Jun–Aug. +
scarious-margined +
connate +  and distinct +
weakly +  and flat +
glabrous +, glabrate +, woolly +  and tomentose +
5 +  and 13 +
Univ. Calif. Publ. Bot. +
pitted +  and smooth +
convex;conic +
6 +  and 12 +
lacerate +, cuneate +  and lanceolate or lance-linear +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
exalbuminous +
erect +  and decumbent +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Eriophyllum lanatum var. cuneatum +  and Eriophyllum lanatum var. monoense +
Eriophyllum lanatum var. integrifolium +
Eriophyllum lanatum +
variety +
campanulate +
plane +, revolute +, entire +  and toothed +
10 cm100 mm <br />0.1 m <br /> (100 cm1,000 mm <br />1 m <br />) +