Erythranthe washingtonensis

(Gandoger) G. L. Nesom

Phytoneuron 2012-39: 39. 2012.

Common names: John Day or Washington monkeyflower
Endemic
Basionym: Mimulus washingtonensis Gandoger Bull. Soc. Bot. France 66: 218. 1919
Treatment appears in FNA Volume 17. Treatment on page 396. Mentioned on page 381.

Annuals, fibrous-rooted or filiform-taprooted. Stems erect to ascending, straight or geniculate at nodes, usually many-branched, terete, 5–25 cm, moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. Leaves cauline, basal not persistent; petiole 2–14 mm; blade palmately veined, deltate or ovate to ovatelanceolate, 4–16 (–23) × 2–11 (–16) mm, base rounded to cuneate or truncate, margins denticulate or entire, apex acute, surfaces moderately puberulent-glandular to villous-glandular, hairs 0.1–0.8 mm, flattened, sometimes vitreous, distinctly multicellular, gland-tipped. Flowers herkogamous, 1–6, from proximal to distal nodes. Fruiting pedicels divergent at nearly right angles, 20–50 mm, densely, minutely stipitate-glandular. Fruiting calyces greenish, ridge-angled, tubular, weakly inflated, 6–8 mm, margins distinctly toothed or lobed, densely, minutely stipitate-glandular, lobes pronounced, erect. Corollas yellow with small reddish-brown dots, abaxial limb with 2 white patches (abaxial ridges), bilaterally symmetric, strongly bilabiate; tube-throat funnelform, 8–10 mm, exserted beyond calyx margin; limb expanded 7–10 mm, lobes obovate-oblong, apex rounded to rounded-cuneate. Styles hispid-hirtellous. Anthers included, glabrous. Capsules included, 5–8.5 mm. 2n = 32.


Phenology: Flowering May–Sep.
Habitat: Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps.
Elevation: 700–1300 m.

Discussion

Erythranthe washingtonensis is considered to be extirpated in Washington by the Washington Natural Heritage Program.

Selected References

None.

Lower Taxa

None.
stipitate-glandular +
2 cm20 mm <br />0.02 m <br /> (5 cm50 mm <br />0.05 m <br />) +
loculicidal +
rounded +  and rounded-cuneate +
Guy L. Nesom +  and Naomi S. Fraga +
(Gandoger) G. L. Nesom +
rosulate +  and subrosulate +
not persistent +
rounded;cuneate or truncate +
Mimulus washingtonensis +
1.6 cm16 mm <br />0.016 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
ovate;ovatelanceolate +
not leathery +  and semifleshy +
0.2 cm2 mm <br />0.002 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
compressed +, inflated +  and tubular +
0.5 cm5 mm <br />0.005 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
John Day or Washington monkeyflower +
regular +  and bilabiate +
yellow with small reddish-brown dots +
compressed +
700–1300 m. +
hairy +  and glabrous +
nodding;spreading;nodding;spreading;erect +
reduced +
Shallow basalt gravel in narrow channels and intermittent streams, sandy stream banks, open slopes, rocky shelves near seeps. +
gland-tipped +  and multicellular +
flattened +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
alternate +, opposite +, rosulate +  and subrosulate +
persistent +  and deciduous +
expanded +
pronounced +
3 (?) +  and Not 5 (?) +
obovate-oblong +
reduced +  and longer +
stipitate-glandular +
lobed +, toothed +, entire +  and denticulate +
basal +  and superior +
tenuinucellate +  and unitegmic +
orthotropous +  and anatropous +
0.2 cm2 mm <br />0.002 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Flowering May–Sep. +
Phytoneuron +
flattened +  and ellipsoid +
3 +  and 5 +
proximal +  and middle +
many-branched +
moderately puberulent-glandular +  and villous-glandular +
straight +
erect;ascending +
hairy +  and glandular-hairy +
terete +  and geniculate +
5 cm50 mm <br />0.05 m <br /> (25 cm250 mm <br />0.25 m <br />) +
hispid-hirtellous +
moderately puberulent-glandular +  and villous-glandular +
Mimulus sect. Erythranthe +  and Mimulus subg. Synplacus +
Erythranthe washingtonensis +
Erythranthe +
species +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
filiform-taprooted +  and fibrous-rooted +
inflated +, tubular +  and ridge-angled +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
perennial +  and annual +
herb +  and subshrub +
terrestrial +  and aquatic +
semiaquatic +  and terrestrial +
9 +, 8 +  and 7 +