Eucladium

Bruch & Schimper

Bryol. Europ. 1: 93. 1846 ,.

Etymology: Greek eu-, good or well, and klados, branch, alluding to well-developed whorls of stem leaves
Treatment appears in FNA Volume 27. Treatment on page 486. Mentioned on page 478, 487, 488, 578.

Plants small, in dense to deep tufts, turfs or swollen cushions, tufa-forming, bright to dark blue-green distally and pale green to yellowish-brown proximally or sometimes throughout, rarely rufous; frequently bleached or whitened and indurated with encrusted lime. Stems slender, branching irregularly, densely foliate in whorls at successive innovations; transverse-section rounded-pentagonal with an irregularly inflated or enlarged hyalodermis and weakly developed sclerodermis; central strand absent; weakly radiculose, rarely strongly so; axillary hairs of 5–10 cells, hyaline throughout. Leaves appressed and scarcely incurved-contorted when dry, erect-spreading when moist, oblong to mostly linear-lanceolate or linear-subulate, base undifferentiated in shape to occasionally broadened-oblong; margins plane, entire except for distinctive irregular hollow teeth on shoulder just distal to the base, not bordered; apex broadly to narrowly acute or subulate; costa strong, percurrent to shortly excurrent as a stout mucro, in transverse-section elliptic; adaxial surface cells quadrate to elongate, adaxial epidermis present, adaxial stereid band, medial guide cells and abaxial stereid band present, abaxial epidermis sometimes absent, hydroid strand absent; basal-cells sharply differentiated across to the margin where they become narrower, elongate, bulging-rectangular to rhomboidal, hyaline, smooth, lax and thin-walled; distal laminal cells subquadrate, walls sometimes irregularly thickened, papillae low, indistinct, simple or 2-fid, several per cell. Specialized asexual reproduction reportedly by rhizoidal brood bodies. Sexual condition dioicous; perigonia terminal, interior leaves longlanceolate, similar to those of the stem; perichaetia terminal, leaves as in stem-leaves, ovatelanceolate, to 2.5 mm and sheathing. Seta to 8 mm, single, erect. Capsule erect, symmetrical, yellowbrown, ovoid to cylindrical; annulus weakly developed, of ca. 2 rows of subvesiculose cells; operculum obliquely conic-rostrate; peristome teeth 16, straight or slightly oblique, lanceolate, finely papillose, entire to variously cleft. Calyptra cucullate, smooth. Spores 8–14 µm, spherical, mostly smooth. Laminal KOH color reaction yellow.

Distribution

North America, Mexico, Central America, Europe, Asia, Africa

Discussion

Species 1.

R. H. Zander (1993) removed Eucladium irroratum (Mitten) Jaeger to the genus Tetracoscinodon, rendering Eucladium monotypic and characterized by absence of stem central strand and sclerodermis, a somewhat enlarged hyalodermis, costa broad in relation to the leaf, plane margins with their peculiar dentition at the leaf shoulder (a condition that may be variable or rarely even absent), and terminal inflorescences. The generic placement is near Trichostomum (Zander). The genus is differentiated from other members of the former tribe Pleuroweisieae by the presence of a peristome, the distinctive serrulations on the proximal leaf margins, and the hyaline, thin-walled and bulging-rectangular differentiated basal cells (Zander 1977).

... more about "Eucladium"
elongate +  and quadrate +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Subulate (?) +  and Acute (?) +
Patricia M. Eckel +
Bruch & Schimper +
reniform +  and rounded +
papillose +  and smooth +
usually slightly wider +
broadened-oblong;undifferentiated +
Undefined subfam. Trichostomaceae +
obovoid +
cucullate +
cleistocarpous +  and stegocarpous +
ovoid;cylindrical +
monoicous +, dioicous +  and sexual +
flat to bulging +
vesiculose +  and thick-walled +
5 (?) +  and 10 (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
percurrent +  and shortly excurrent +
North America +, Mexico +, Central America +, Europe +, Asia +  and Africa +
differentiated +
Greek eu-, good or well, and klados, branch, alluding to well-developed whorls of stem leaves +
multicellular +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
incurved-contorted +
ovatelanceolate +, oblong +  and mostly linear-lanceolate or linear-subulate +
reduced +
0 cm0 mm <br />0 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
encrusted +
indurate +
bulging-rectangular +  and rhomboidal +
narrower +
conic-rostrate +
photosynthetic +
dioicous +  and sexual +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Bryol. Europ. +
osada1958a +  and saito1972a +
straight +
elongate +
0 cm0 mm <br />0 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
2 +  and 1 +
yellow +  and laminal +
spherical +
foliate +  and branching +
rounded-triangular +  and rounded-pentagonal +
slender +
0.2 cm2 mm <br />0.002 m <br /> (4 cm40 mm <br />0.04 m <br />) +
quadrate;elongate +
Eucladium +
Pottiaceae subfam. Trichostomoideae +
ovoid;cylindric +
not bordered +  and hollow +
distinctive +
orange +  and yellow +
stemless +  and perigoniate +
whitened;rufous;dark blue-green distally and pale green;yellowish-brown +