Euphorbia brachycera

Engelmann in W. H. Emory

Rep. U.S. Mex. Bound. 2(1): 192. 1859.

Common names: Horned or shorthorn spurge
Synonyms: Euphorbia brachycera var. robusta (Engelmann) Dorn E. montana var. robusta Engelmann E. odontadenia Boissier E. robusta (Engelmann) Small Tithymalus brachycerus (Engelmann) Small T. robustus (Engelmann) Small
Treatment appears in FNA Volume 12. Treatment on page 300. Mentioned on page 296, 305, 313.

Herbs, perennial, with thick rootstock. Stems erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. Leaves: petiole 0–0.5 mm; blade oblongelliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. Cyathial arrangement: terminal pleiochasial branches 3–5 (–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. Cyathia: peduncle 0.4–1 mm. Involucre turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. Staminate flowers 10–15. Pistillate flower: ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. Capsules depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. Seeds light gray, cylindric ovoid-oblong, 2–2.8 (–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. 2n = 28.


Phenology: Flowering and fruiting spring–summer.
Habitat: Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests.
Elevation: 1200–3200 m.

Distribution

V12 661-distribution-map.jpg

Ariz., Colo., Idaho, Mont., Nebr., N.Mex., N.Dak., S.Dak., Tex., Utah, Wyo., Mexico (Chihuahua), Mexico (Coahuila), Mexico (Sonora)

Discussion

Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque.

Selected References

None.

Lower Taxa

None.

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... more about "Euphorbia brachycera"
mucronate;obtuse;mucronate;obtuse;acute +
Ricarda Riina +, Dmitry V. Geltman +, Jess A. Peirson +  and Paul E. Berry +
Engelmann in W. H. Emory +
obtuse +, attenuate +, acute +, rounded +  and truncate +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
oblanceolate;broadly ovate +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dichasial +  and pleiochasial +
distinct +
rhombic +  and triangular-ovate +
cymose +, branched +, unbranched +  and pleiochasial +
arranged +  and opposite +
additional;2;4 +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
3-lobed +  and depressed-ovoid +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
sessile +  and shortly stipitate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
puberulent +  and glabrous +
rounded +
0.24 cm2.4 mm <br />0.0024 m <br /> (0.33 cm3.3 mm <br />0.0033 m <br />) +
Horned or shorthorn spurge +
Ariz. +, Colo. +, Idaho +, Mont. +, Nebr. +, N.Mex. +, N.Dak. +, S.Dak. +, Tex. +, Utah +, Wyo. +, Mexico (Chihuahua) +, Mexico (Coahuila) +  and Mexico (Sonora) +
1200–3200 m. +
pistillate +  and staminate +
puberulent +  and glabrous +
0.5mm;0.8mm +
subtending +
crescent-shaped +  and triangular-ovate +
0.7mm +  and 1.7mm +
Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
axillary +  and terminal +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
puberulent +  and glabrous +
cupulate +, campanulate +  and turbinate +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
caducous +, deciduous +  and persistent +
inconspicuous +
bractlike +
palmate +  and pinnate +
occasionally slightly crenate +  and dentate +
hairy;glabrous +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
connate;distinct +
hairy +  and glabrous +
0 cm0 mm <br />0 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
Flowering and fruiting spring–summer. +
Rep. U.S. Mex. Bound. +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
light gray +
0.2 cm2 mm <br />0.002 m <br /> (0.28 cm2.8 mm <br />0.0028 m <br />) +
ovoid-oblong +  and cylindric +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
free +, connate +  and distinct +
ascending +  and erect +
puberulent +  and glabrous +
10 cm100 mm <br />0.1 m <br /> (60 cm600 mm <br />0.6 m <br />) +
herbaceous +  and woody +
deciduous +  and persistent +
connate +  and distinct +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (?) +  and 0.09 cm0.9 mm <br />9.0e-4 m <br /> (?) +
puberulent +  and glabrous +
Euphorbia brachycera var. robusta +, E. montana var. robusta +, E. odontadenia +, E. robusta +, Tithymalus brachycerus +  and T. robustus +
Euphorbia brachycera +
Euphorbia subg. Esula +
species +
1-4+ times 2-branched +  and pleiochasial +
rounded +  and attenuate +
10 +, 9 +, 8 +, 7 +  and 6 +