Euphorbia perennans

(Shinners) Warnock & M. C. Johnston

SouthW. Naturalist 5: 170. 1960.

Common names: Terlingua spurge perennial sandmat
Basionym: Chamaesyce perennans Shinners Field & Lab. 24: 38. 1956
Treatment appears in FNA Volume 12. Treatment on page 282. Mentioned on page 257, 270.

Herbs, perennial, with strongly thickened, woody rootstock. Stems erect, 7–45 cm, glabrous. Leaves opposite; stipules distinct, linear-filiform in (1–) 2 (–3) segments, 0.3–0.4 mm, glabrous; petiole 0.8–2 mm, glabrous; blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous. Cyathia solitary at distal nodes or at nodes of short, axillary branches; peduncle 1.8–3 mm. Involucre broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous; glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm; appendages absent. Staminate flowers 35–45. Pistillate flowers: ovary glabrous; styles 0.7–0.9 mm, 2-fid nearly entire length. Capsules subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous; columella 2.2–2.7 mm. Seeds white to light-brown, ovoid, 3–4-angled in cross-section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled.


Phenology: Flowering and fruiting spring–fall.
Habitat: Desert scrub, on cretaceous and gypseous clay, limestone hills and flats.
Elevation: 900–1200 m.

Discussion

Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011).

Euphorbia perennans is known in the flora area only from Brewster County.

Selected References

None.

Lower Taxa

None.

"connate" is not a number. "distinct" is not a number."entire" is not a number.

... more about "Euphorbia perennans"
acute;rounded +
Victor W. Steinmann +, Jeffery J. Morawetz +, Paul E. Berry +, Jess A. Peirson +  and Ya Yang +
(Shinners) Warnock & M. C. Johnston +
pinnate +, symmetric +  and asymmetric +
rounded +  and cordate +
Chamaesyce perennans +
0.5 cm5 mm <br />0.005 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
orbiculate-deltate;reniform-deltate +
0.4 cm4 mm <br />0.004 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.33 cm3.3 mm <br />0.0033 m <br />) +
subglobose +  and broadly ovoid +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
Terlingua spurge +  and perennial sandmat +
Tex. +  and Mexico (Chihuahua) +
900–1200 m. +
pistillate +  and staminate +
green +  and yellow-green +
0.3mm;0.5mm +
subtending +
elliptic +  and oblong +
0.7mm +  and 1.4mm +
Desert scrub, on cretaceous and gypseous clay, limestone hills and flats. +
axillary +  and terminal +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
broadly-campanulate +  and hemispheric +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
caducous +, deciduous +  and persistent +
bractlike +
palmate +  and pinnate +
toothed +  and entire +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
connate;distinct +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering and fruiting spring–fall. +
SouthW. Naturalist +
white;light-brown +
0.2 cm2 mm <br />0.002 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
smooth +  and faintly transverse-wrinkled +
3-4-angled +  and ovoid +
0.1 cm1 mm <br />0.001 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
free +, connate +  and distinct +
flattened +  and terete +
7 cm70 mm <br />0.07 m <br /> (45 cm450 mm <br />0.45 m <br />) +
interpetiolar +
linear-filiform +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
connate +  and distinct +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (?) +  and 0.09 cm0.9 mm <br />9.0e-4 m <br /> (?) +
glaucous +  and glabrous +
Chamaesyce +
Euphorbia perennans +
Euphorbia sect. Anisophyllum +
species +
axillary +  and terminal +
10 +, 9 +, 8 +, 7 +  and 6 +