Frangula rubra

(Greene) Grubov

Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 8: 271. 1949.

Common names: Sierra coffeeberry
Endemic
Basionym: Rhamnus rubra Greene Pittonia 1: 68. 1887
Treatment appears in FNA Volume 12. Treatment on page 55. Mentioned on page 53, 56, 58.

Shrubs, 0.5–2 m. Stems red to gray, glabrous or hairy. Leaves deciduous; petiole 2–12 mm; blade light to bright green abaxially, green or dull green adaxially, narrowly elliptic to oblong or obovate, 1.5–8.5 cm, ± herbaceous, base rounded to obtuse or acute, margins entire or serrulate to denticulate, apex acute to obtuse or rounded, surfaces glabrous or short-puberulent, or abaxial puberulent on midrib and veins; secondary-veins (7–) 8–11 pairs. Inflorescences umbels, pedunculate, (2–) 4–15-flowered. Pedicels 1–12 mm. Stigmas 2-lobed. Drupes black, globose or pyriform, 8–12 mm; stones 2 (–3).

Discussion

Subspecies 5 (5 in the flora).

The Frangula rubra complex is a group of closely related populations that needs study. In Nevada, they occur only in Douglas and Washoe counties. Descriptions by C. B. Wolf (1938) provide only a single distinct character to separate the subspecies, and he reported much intergradations between subsp. obtusissima and all the other subspecies. Field and herbarium studies argue for the recognition of infraspecific taxa despite the intermediates.

Subspecies yosemitana and Frangula californica subsp. cuspidata grow along the east side of the Sierra Nevada and can be easily confused. Plants of both taxa can be deciduous, but Wolf noted differences in pubescence and leaf margin to differentiate the two, although both are variable throughout their ranges.

Selected References

None.

Key

1 Leaf blade surfaces short-puberulent. Frangula rubra subsp. yosemitana
1 Leaf blade surfaces glabrous or abaxial puberulent on midrib and veins. > 2
2 Young branches gray; leaves clustered on short-shoots. Frangula rubra subsp. modocensis
2 Young branches usually red, rarely gray; leaves scattered along branchlets. > 3
3 Leaf blade bases and apices rounded. Frangula rubra subsp. obtusissima
3 Leaf blade bases and apices acute to obtuse. > 4
4 Leaf blades bright green; drupes globose. Frangula rubra subsp. rubra
4 Leaf blades usually dull green, especially abaxially; drupes pyriform. Frangula rubra subsp. nevadensis
... more about "Frangula rubra"
perigynous +  and epigynous +
acute +  and obtuse or rounded +
John O. Sawyer† +  and Guy L. Nesom +
(Greene) Grubov +
rounded +  and obtuse or acute +
Rhamnus rubra +
protruding +
light +  and bright green abaxially +
3[-5]-veined +  and pinnate +
spinulose +, spinose +, serrate +  and entire +
denticulate +, dentate +, crenulate +, crenate +  and serrulate +
narrowly elliptic;oblong or obovate +
1.5 cm15 mm <br />0.015 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
Sierra coffeeberry +
Calif. +, Nev. +  and Oreg. +
pyriform +  and globose +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
circumscissile +
free +  and adnate +
unisexual +  and bisexual +
(2-)4-15-flowered +  and pedunculate +
deciduous +
serrulate to denticulate +  and entire +
intrastaminal +
0.1 cm1 mm <br />0.001 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
not fleshy +
perigynous +  and epigynous +
adnate +  and distinct +
clawed +, broadly obovate +  and obcordate +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
2-4-carpellate +
Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. +
not furrowed +
obovoid +  and lenticular +
yellowish +  and green or white +
distinct +
spreading +  and erect +
keeled;ovate-triangular +
red +  and gray +
hairy +  and glabrous +
indehiscent +
short-puberulent +  and glabrous +
Rhamnus sect. Frangula +  and Rhamnus subg. Frangula +
Frangula rubra +
Frangula +
species +
deciduous +  and evergreen +
polygamous +, dioecious +  and synoecious +