Grimmia anomala

Hampe in W. P. Schimper

in W. P. Schimper, Syn. Musc. Eur. ed. 2, 270. 1876,.

Synonyms: Grimmia hartmanii subsp. anomala (Hampe) Loeske Grimmia pachyneurula Britton Grimmia philibertiana
Treatment appears in FNA Volume 27. Treatment on page 248. Mentioned on page 227, 247, 251, 258.

Plants in tufts, yellowish green, blackish proximally. Stems 1.5–3.5 cm, small central strand present. Leaves irregularly imbricate when dry, erect when moist, oblong-lanceolate, gradually narrowed into a blunt chlorophyllose point, 1.5–2.5 × 0.4–0.8 mm, keeled, margins recurved on one or both sides, awns absent to very short, costa projecting on abaxial side; basal juxtacostal laminal cells quadrate to short-rectangular, straight, thin-walled; basal marginal laminal cells quadrate to short-rectangular with thickened transverse walls; medial laminal cells roundedquadrate, walls slightly sinuose, thin or thick-walled; distal laminal cells 1-stratose, in places 2-stratose, margins 1-stratose. Gemmae in clusters, globular, yellowish green to orange, multicellular, on hyaline, deformed leaf apices. Sexual condition dioicous. Seta straight to slightly arcuate when moist, 3–5 mm. Capsule extremely rare, exserted, brownish, oblong-ovoid, smooth, exothecial cells isodiametric, thick-walled, annulus present, operculum with long straight beak, peristome teeth orange, fully-developed, smooth proximally, perforated and papillose distally. Calyptra mitrate.


Habitat: Exposed, damp acidic rock in boreal and alpine meadows and slopes
Elevation: moderate to high elevations (200-3000 m)

Distribution

V27 341-distribution-map.gif

Greenland, Alta., B.C., Nfld. and Labr. (Nfld.), N.S., Ont., Alaska, Calif., Colo., Idaho, Mich., Mont., Oreg., Utah, Wash., Wyo., Eurasia

Discussion

Because of the always abundant gemmae on leaf apices, Grimmia anomala has often been confused with G. hartmanii. Indeed, it has frequently been treated as a variety of that species (G. N. Jones 1933; E. Lawton 1971). However, as pointed out by O. Vitikainen (1969), the ecological and morphological features of G. hartmanii are so constant and distinct that there is no doubt that it can be treated as a separate species. Special features of G. anomala are stems with central strands, and longitudinal ridges on cell walls in the distal part of the leaf that resemble papillae in transverse section. It tends to prefer upland habitats. In contrast, G. hartmanii is principally a lowland species, frequently growing on boulders in forests. It lacks a central strand, and longitudinal ridges are absent. Its spreading, secund distal leaves taper to long, sharply keeled apices.

Selected References

None.

Lower Taxa

None.
... more about "Grimmia anomala"
deciduous +  and persistent +
well-defined +
differentiated +
Roxanne I. Hastings +  and Henk C. Greven +
Hampe in W. P. Schimper +
only rarely longer +
not papillose +
toothed +
sinuous +  and straight to sometimes +
sinuose-nodulose;quadrate;elongate +
oblate;elongate +
divided +
straight +
undifferentiated +
plicate +  and smooth +
less +  and 1/2 +
not papillose +
small;medium +
pendulous +  and cernuous +
striate +  and smooth +
oblong-ovoid +
thin +  and thick +
thick-walled +  and thin-walled +
quadrate +  and rectangular +
excurrent;percurrent;excurrent +
subpercurrent +
reniform +, elliptical +  and terete +
usually not markedly larger +
1-stratose +  and multistratose +
2-stratose +  and 1-stratose +
Greenland +, Alta. +, B.C. +, Nfld. and Labr. (Nfld.) +, N.S. +, Ont. +, Alaska +, Calif. +, Colo. +, Idaho +, Mich. +, Mont. +, Oreg. +, Utah +, Wash. +, Wyo. +  and Eurasia +
moderate to high elevations (200-3000 m) +
reproduction +  and asexual +
dioicous +  and sexual +
Exposed, damp acidic rock in boreal and alpine meadows and slopes +
papillose +, mammillose +  and smooth +
specialized +
dioicous +, autoicous +  and sexual +
1.5mm;2.5mm +
keeled +, narrowed +  and oblong-lanceolate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
thick-walled +  and sinuose +
thick-walled +  and sinuose +
rostrate;conic;rostrate;conic;mammillate +
not +  and enlarged +
not +  and differentiated +
lanceolate;linear +
in W. P. Schimper, Syn. Musc. Eur. ed. +
straight +  and slightly arcuate +
sigmoid +  and arcuate +
papillose +  and smooth +
arcuate;cygneous +
short to long +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
ornamented +  and smooth +
globose +
1.5 cm15 mm <br />0.015 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
muticous;long-awned +
Grimmia hartmanii subsp. anomala +, Grimmia pachyneurula +  and Grimmia philibertiana +
Grimmia anomala +
Grimmia subg. Rhabdogrimmia +
species +
thick-walled +  and sinuose +
blackish;yellowish green +