Grimmia ramondii

(Lamarck & De Candolle) Margadant

Lindbergia 1: 128. 1972,.

Basionym: Pterigynandrum ramondii Lamarck & de Candolle Fl. Franç. ed. 3, 2: 462. 1805 Grimmia curvata (Bridel) De Sloover G. serrata Kindberg
Synonyms: Dryptodon patens (Hedwig) Bridel Racomitrium patens (Hedwig) Huebener
Treatment appears in FNA Volume 27. Treatment on page 256. Mentioned on page 227.

Plants in robust, loose, arched-ascending, readily disintegrating patches, brownish green distally, blackish proximally. Stems 5–10 cm, central strand absent. Leaves loosely appressed when dry, patent when moist, broadly ovatelanceolate, tapering to an acute, slightly toothed apex, 2–3 × 0.4–0.6 mm, keeled, margins recurved on both sides, awns absent, costa stout, with two abaxial lamellae (or wings) forming parallel ridges along its length; basal juxtacostal laminal cells yellow, linear, sinuose, thick-walled; basal marginal laminal cells quadrate to short-rectangular, slightly thick-walled; medial laminal cells quadrate to rectangular, sinuous, thick-walled; distal laminal cells 1-stratose, margins 2-stratose. Gemmae absent. Sexual condition dioicous. Seta cygneous to arcuate at maturity, flexuose when old, 3–5 mm. Capsule occasionally present, exserted, obloid, yellowish green to yellowish-brown, plicate when empty, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth purple, divided nearly to base into two partly adhering segments, papillose. Calyptra mitrate.


Habitat: Dry to damp acidic rock
Elevation: low to high elevations (0-2000 m)

Distribution

V27 354-distribution-map.gif

Greenland, Alta., B.C., Nfld. and Labr. (Nfld.), N.W.T., Alaska, Calif., Idaho, Mich., Mont., Oreg., Wash., Europe, Asia (China), Asia (Japan), n Africa

Discussion

Grimmia ramondii occurs near sea level in arctic areas and along the West Coast. In the latter region, however, it is more common in mid-elevation montane sites and may also be found above the tree line. It differs from most Grimmia species by its lack of awns, winged costa, and tall, loose growth form. At first sight, G. ramondii has the habit of a Racomitrium. However, the sporophyte has an arcuate rather than straight seta, and the capsules are striate rather than smooth. Because of its habit and the absence of a central strand, K. G. Limpricht ([1885–]1890–1903, vol. 1) placed it, together with G. hartmanii and G. atrata, in Dryptodon, a genus intermediate between Grimmia and Racomitrium. There are many other Grimmia species, however, without a central strand, and G. hartmanii and G. atrata have already been removed from Dryptodon, so there is no reason to maintain that monotypic genus. Grimmia ramondii may be confused with G. hartmanii and G. elatior. However, the lack of awns, the Racomitrioideae-like areolation, and the distinct costal wings distinguish it easily from those species.

Selected References

None.

Lower Taxa

None.
... more about "Grimmia ramondii"
deciduous +  and persistent +
well-defined +
differentiated +
Toothed (?) +  and Acute (?) +
Roxanne I. Hastings +  and Henk C. Greven +
(Lamarck & De Candolle) Margadant +
only rarely longer +
not papillose +
toothed +
sinuous +  and straight to sometimes +
sinuose-nodulose;quadrate;elongate +
oblate;elongate +
divided +
Pterigynandrum ramondii +  and Grimmia curvata +
undifferentiated +
plicate +  and smooth +
less +  and 1/2 +
not papillose +
small;medium +
yellowish green +  and yellowish-brown +
pendulous +  and cernuous +
striate +  and smooth +
thin +  and thick +
thick-walled +  and thin-walled +
quadrate +  and rectangular +
excurrent;percurrent;excurrent +
subpercurrent +
reniform +, elliptical +  and terete +
usually not markedly larger +
1-stratose +  and multistratose +
Greenland +, Alta. +, B.C. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, Alaska +, Calif. +, Idaho +, Mich. +, Mont. +, Oreg. +, Wash. +, Europe +, Asia (China) +, Asia (Japan) +  and n Africa +
low to high elevations (0-2000 m) +
reproduction +  and asexual +
dioicous +  and sexual +
Dry to damp acidic rock +
papillose +, mammillose +  and smooth +
thick-walled +  and sinuose +
specialized +
dioicous +, autoicous +  and sexual +
2mm;3mm +
keeled +, tapering +  and ovatelanceolate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
quadrate +  and rectangular +
thick-walled +  and sinuose +
thick-walled +  and sinuose +
disintegrating +
arched-ascending +
not +  and enlarged +
not +  and differentiated +
lanceolate;linear +
flexuose +
sigmoid +  and arcuate +
papillose +  and smooth +
cygneous;arcuate +
short to long +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
ornamented +  and smooth +
globose +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
muticous;long-awned +
Dryptodon patens +  and Racomitrium patens +
Grimmia ramondii +
Grimmia subg. Rhabdogrimmia +
species +
blackish;brownish green +