Grusonia clavata

(Engelmann) H. Robinson

Phytologia 26: 176. 1973.

Common names: Club-c holla
IllustratedEndemic
Basionyms: Opuntia clavata Engelmann in F. A. Wislizenus, Mem. Tour N. Mexico, 95. 1848
Synonyms: Cactus clavatus (Engelmann) Lemaire Corynopuntia clavata (Engelmann) F. M. Knuth
Treatment appears in FNA Volume 4. Treatment on page 121. Mentioned on page 119, 120.

Shrubs, forming mats, widely spreading, 5–15 cm, in series of usually 1–2 stem segments. Roots diffuse. Stem segments short-clavate, strongly narrowed at base, 2.5–5 (–7.5) × 1.5–3 cm; tubercles prominent, oval, 5–10 (–16) mm, narrow, 4–6 times longer than wide, obscured by interlacing spines; areoles 3 mm in diam.; wool white to gray. Spines 7–15 per areole, primarily in distal areoles; major 3–5 abaxial spines deflexed, white, flattened, longest central spine daggerlike, broadly tapered, 12–35 × 1.5+ mm basally; major 1–3 adaxial spines ascending, white, yellow to tan, angular-flattened to subterete. Glochids in adaxial 1/4 of areole, yellowish white, ± 4 mm. Flowers: inner tepals bright-yellow, to 25 mm; filaments light yellow-green to white; style and stigma lobes white. Fruits yellow, barrel-shaped to ellipsoid, 30–45 × 15–25 mm, fleshy, spineless, densely ± yellow-glochidiate; areoles 35–55. Seeds yellowish white, 4.5 × 4 mm, smooth. 2n = 22.


Phenology: Flowering late spring (May–Jun).
Habitat: Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils
Elevation: 1800-2500 m

Discussion

L. D. Benson (1982) mapped “Opuntia clavata” in northeast Arizona, but no supporting specimens have been found. The report may be based on low-growing specimens of Cylindropuntia whipplei, a cholla common in that area.

Selected References

None.

Lower Taxa

None.

... more about "Grusonia clavata"
yellow +  and tan +
cushionlike +
3 cm30 mm <br />0.03 m <br /> (?) +
0.1 cm1 mm <br />0.001 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
deciduous +  and persistent +
35 +  and 55 +
cushionlike +
Donald J. Pinkava +
(Engelmann) H. Robinson +
curving +
dislodged +
cylindric-clavate +
Opuntia clavata +
proliferating +
flattened +  and cylindric +
Club-c holla +
1800-2500 m +
light yellow-green +  and white +
15 cm150 mm <br />0.15 m <br /> (30 cm300 mm <br />0.3 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (15 cm150 mm <br />0.15 m <br />) +
indehiscent +
persistent +, long +  and deciduous +
30mm;45mm +
yellow-glochidiate +
barrel-shaped +  and ellipsoid +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
not protruding +
4 cm40 mm <br />0.04 m <br /> (?) +
Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils +
0 cm0 mm <br />0 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
deciduous +
flattened +  and conic +
0 cm0 mm <br />0 m <br /> (3 cm30 mm <br />0.03 m <br />) +
succulent +
tuberculate +
fluted;spheric;depressed-spheric or club-shaped +
1.2 cm12 mm <br />0.012 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
tapered +  and daggerlike +
sometimes covered by outgrowths of proximal portion of style base or of flower tube wall +  and open +
persistent +  and deciduous +
Flowering late spring (May–Jun). +
ridgelike +  and nipple--shaped +
3;2;2;30 +
adventitious +
tuberlike +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
4.5 cm45 mm <br />0.045 m <br /> (?) +
pubescent +  and glabrous +
flattened;suborbicular +
4 cm40 mm <br />0.04 m <br /> (?) +
Illustrated +  and Endemic +
interlacing +
deciduous +  and persistent +
conspicuous +
7 (?) +  and 15 (?) +
terete +  and strongly flattened +
hard +  and rigid +
decurrent +
winged +  and unsegmented +
tuberculate +  and smooth +
5 cm50 mm <br />0.05 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
2.5 cm25 mm <br />0.025 m <br /> (5 cm50 mm <br />0.05 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (3 cm30 mm <br />0.03 m <br />) +
segmented +
woody +  and succulent +
Cactus clavatus +  and Corynopuntia clavata +
adventitious +
tuberlike +
epipetric +  and epiphytic +
Grusonia clavata +
Grusonia +
species +
dull +  and glossy +
1 cm10 mm <br />0.01 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
obscured +  and prominent +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
conspicuous +
terrestrial;epiphytic or epipetric +
white +  and gray +