Gymnocarpium disjunctum

(Ruprecht) Ching

Acta Phytotax. Sin. 10: 304. 1965.

Common names: Western oak fern
Basionym: Polypodium dryopteris var. disjunctum Ruprecht Distr. Crypt. Vasc. Ross., 52. 1845
Synonyms: Dryopteris disjuncta (Ruprecht) C. V. Morton Gymnocarpium dryopteris subsp. disjunctum (Ruprecht) Sarvela Gymnocarpium dryopteris var. disjunctum (Ruprecht) Ching
Treatment appears in FNA Volume 2.

Stems 1–3 mm diam.; scales 2–4 mm. Fertile leaves usually 20–68 cm. Petiole 12–44 cm with sparse glandular-hairs distally; scales 2–6 mm. Blade broadly deltate, 3-pinnate-pinnatifid, 8–24 cm, lax and delicate, abaxial surface and rachis glabrous or with sparse glandular-hairs, adaxial surface glabrous. Pinna apex acuminate. Proximal pinnae 5–18 cm, ± perpendicular to rachis, with basiscopic pinnules ± perpendicular to costa; basal basiscopic pinnule sessile, pinnate-pinnatifid (with basal pinnulets, and sometimes 2 adjacent pinnulets, separate), basal basiscopic pinnulet usually longer (sometimes equaling or shorter) than adjacent pinnulet; 2d basal basiscopic pinnule sessile with basal basiscopic pinnulet usually longer than or equaling adjacent pinnulet; basal acroscopic pinnule sessile, with basal basiscopic pinnulet usually longer than or equaling adjacent pinnulet. Pinnae of 2d pair usually sessile with basal basiscopic pinnule longer than or equaling adjacent pinnule and markedly longer than basal acroscopic pinnule; basal acroscopic pinnule distinctly shorter than adjacent pinnule or rarely absent, apex often crenulate, obtuse. Pinnae of 3d pair usually sessile with basal basiscopic pinnule longer than or equaling adjacent pinnule and longer than basal acroscopic pinnule; basal acroscopic pinnule shorter than adjacent pinnule. Ultimate segments of proximal pinnae oblong, crenate to slightly lobed, apex crenulate, acute. Spores 27–31 µm. 2n = 80.


Habitat: Shaded, rocky slopes and ravines, mixed coniferous woods, moist stream and creek banks
Elevation: 0–2400 m

Distribution

V2 701-distribution-map.gif

Alta., B.C., Alaska, Idaho, Mont., Oreg., Wash., Wyo., Asia in ne former Soviet republics

Discussion

In addition to the west coast of North America, Gymnocarpium disjunctum is found on Sakhalin Island in southern Kamchatka, in the former Soviet republics.

Of conservation concern.

Selected References

None.

Lower Taxa

None.

"-3timeslengthofblade" is not declared as a valid unit of measurement for this property.

... more about "Gymnocarpium disjunctum"
with sparse glandular-hairs +  and glabrous +
acute +, crenulate +  and obtuse +
Kathleen M. Pryer +
(Ruprecht) Ching +
not swollen +
Polypodium dryopteris var. disjunctum +
3-pinnate-pinnatifid;deltate +
reduced +
8 cm80 mm <br />0.08 m <br /> (24 cm240 mm <br />0.24 m <br />) +
roundish +
Western oak fern +
Alta. +, B.C. +, Alaska +, Idaho +, Mont. +, Oreg. +, Wash. +, Wyo. +  and Asia in ne former Soviet republics +
0–2400 m +
often +  and glabrous +
of minute glands +  and lacking +
not continuous +
Shaded, rocky slopes and ravines, mixed coniferous woods, moist stream and creek banks +
monomorphic +
20 cm200 mm <br />0.2 m <br /> (68 cm680 mm <br />0.68 m <br />) +
12 cm120 mm <br />0.12 m <br /> (44 cm440 mm <br />0.44 m <br />) +
shorter +  and distinctly shorter +
longer +  and markedly longer +
acroscopic +, basiscopic +  and perpendicular +
pinnate-pinnatifid +
5 cm50 mm <br />0.05 m <br /> (18 cm180 mm <br />0.18 m <br />) +
Acta Phytotax. Sin. +
persistent +
round +, cuplike +, hoodlike +, reniform +, falcate +  and linear +
2 +  and 3 +
persistent +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
entire +  and crenate +
discrete +
reniform +  and oblong +
dictyostelic +, unbranched +  and branched +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
creeping +  and erect +
Dryopteris disjuncta +, Gymnocarpium dryopteris subsp. disjunctum +  and Gymnocarpium dryopteris var. disjunctum +
Gymnocarpium disjunctum +
Gymnocarpium +
species +
epiphytic +  and hemiepiphytic +