Helenium elegans var. amphibolum

(A. Gray) Bierner

Sida 5: 47. 1972.

Basionym: Helenium amphibolum A. Gray Proc. Amer. Acad. Arts 9: 202. 1874
Treatment appears in FNA Volume 21. Treatment on page 433.

Plants 25–70 cm. Leaves: proximal blades usually irregularly toothed or lobed; mid and distal leaves usually entire. Heads 20–50. Peduncles 3–14 cm. Involucres 7–11 × 8–12 (–13) mm. Ray-florets 13–17; corollas 6.3–15 (–19) × 2.2–5.5 (–8.7) mm. Disc-florets 250–500 (–700+). Cypselae 0.7–1 mm; pappi 0.1–0.3 (–0.4) mm. 2n = 26.


Phenology: Flowering (Feb–)Mar–Jun(–Sep).
Habitat: Calcareous soils, ditches, washes, along streams
Elevation: 300–1900 m

Distribution

V21-1085-distribution-map.gif

Tex., Mexico (Coahuila), Mexico (Nuevo León), Mexico (San Luis Potosí), Mexico (Tamaulipas)

Discussion

Selected References

None.

Lower Taxa

None.
not tailed +, rounded +  and obtuse +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
paniculiform +
indeterminate +  and determinate +
Mark W. Bierner +
(A. Gray) Bierner +
decurrent +
Helenium amphibolum +
compound +  and simple +
pinnatifid;lobed;spatulate;ovate;spatulate;ovate;oblong;oblanceolate;linear;lanceolate;elliptic +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
1.5 cm15 mm <br />0.015 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
0.23 cm2.3 mm <br />0.0023 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.87 cm8.7 mm <br />0.0087 m <br />) +
yellow +  and brown +
0.63 cm6.3 mm <br />0.0063 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
4-5-angled +  and obpyramidal +
fertile +  and bisexual +
fertile +  and bisexual +
lanceolate;linear +
Tex. +, Mexico (Coahuila) +, Mexico (Nuevo León) +, Mexico (San Luis Potosí) +  and Mexico (Tamaulipas) +
300–1900 m +
gland-dotted +
glabrous or +  and sparsely densely hairy +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Calcareous soils, ditches, washes, along streams +
discoid +  and radiate +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
1.2 cm12 mm <br />0.012 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
ovoid;obconic;ovoid;obconic;hemispheric;globose +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
sessile +  and petiolate +
hairy +  and glabrous +
deltate +
dentate to pinnatifid or palmatifid +  and entire +
connate +  and distinct +
ovate +, oblong +, oblanceolate +, linear +, lanceolate +  and elliptic +
unequal +  and subequal +
2-carpellate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
persistent +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
3 cm30 mm <br />0.03 m <br /> (14 cm140 mm <br />0.14 m <br />) +
Flowering (Feb–)Mar–Jun(–Sep). +
9 +  and 34 +
lobed +  and toothed +
bearing subulate enations +, hairy +  and bristly +
ovoid;hemispheric;ovoid;hemispheric;globose;conic +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
6 +  and 7 +
non-aristate +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
exalbuminous +
winged +  and branched +
1 (?) +  and 10 (?) +
glabrous or +  and sparsely moderately hairy +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Helenium elegans var. amphibolum +
Helenium elegans +
variety +
urceolate +  and campanulate +
dilated +
toothed +  and entire +
20 cm200 mm <br />0.2 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +