Heterotheca camporum var. glandulissima

(as glandulissimum) Semple

Brittonia 35: 146, fig. 1. 1983.

Endemic
Synonyms: Chrysopsis camporum var. glandulissima (Semple) Cronquist
Treatment appears in FNA Volume 20. Treatment on page 256. Mentioned on page 255.

Plants 35–140 cm. Distal stems and peduncles moderately to densely stipitate-glandular. Distal leaf faces moderately hispido-strigose, sparsely to moderately stipitate-glandular. 2n = 36.


Phenology: Flowering Jul–Oct(–Dec).
Habitat: Limestone bluffs and glades, stream banks, margins of mixed mesophytic pine-oak woods, fields, roadsides, disturbed sandy areas
Elevation: 30–800+ m

Distribution

V20-557-distribution-map.gif

Ala., Ark., Ky., Mich., Miss., Mo., N.J., N.C., Tenn., Va.

Discussion

Variety glandulissima grows in the Ridge and Valley region of central and eastern Tennessee and adjacent Alabama, Kentucky, Mississippi, and Virginia. It is introduced and established in northern Arkansas, southern Missouri, northeastern North Carolina; occurrences in southern Michigan and northern New Jersey probably were emphemeral and did not persist.

Variety glandulissima often is taller than 100 cm and obviously stipitate-glandular. The variety appears to have evolved within historic times in Tennessee, or only recently to have evolved the traits necessary to make it a successful roadside weed. J. C. Semple (1983) discussed its range expansion over the last 100 years; it continues to expand its range in the eastern United States.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
corymbiform +, open +  and congested +
indeterminate +  and determinate +
John C. Semple +
(as glandulissimum) Semple +
cuneate +
Chrysopsis camporum +
compound +  and simple +
1.5 cm15 mm <br />0.015 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
38 cm380 mm <br />0.38 m <br /> (73 cm730 mm <br />0.73 m <br />) +
ovate;obovate;oblanceolate;oblanceolate +
reduced +
6mm;15mm +
rugulose +  and muricate +
barbellulate +  and barbellate +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.66 cm6.6 mm <br />0.0066 m <br />) +
obconic +  and compressed +
fertile +  and bisexual +
stipitate-glandular +
stipitate-glandular +
Ala. +, Ark. +, Ky. +, Mich. +, Miss. +, Mo. +, N.J. +, N.C. +, Tenn. +  and Va. +
30–800+ m +
strigose +, stipitate-glandular +, moderately strigose +  and sparsely moderately stipitate-glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Limestone bluffs and glades, stream banks, margins of mixed mesophytic pine-oak woods, fields, roadsides, disturbed sandy areas +
paniculiform +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
tan;rusty brown +
attenuate +  and clavate +
0.9 cm9 mm <br />0.009 m <br /> (1.05 cm10.5 mm <br />0.0105 m <br />) +
cylindric;campanulo-hemispheric +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (2.15 cm21.5 mm <br />0.0215 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
10.5mm;17mm +
sessile +  and petiolate +
stipitate-glandular +  and hispido-strigose +
cauline +  and basal +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
fimbriate-ciliate +  and strigoso-ciliate +
stipitate-glandular +  and strigose +
2-carpellate +
in 3 +  and 4 series +
stipitate-glandular +  and short-hispido-strigose +
0.5 cm5 mm <br />0.005 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
Flowering Jul–Oct(–Dec). +
lanceolate +  and linear-lanceolate +
ovate-oblanceolate +
stipitate-glandular +  and hairy +
linear-lanceolate +
subpetiolate +  and sessile +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
7 +  and 10 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
ascending +  and erect +
1 +  and 30 +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis camporum var. glandulissima +
Heterotheca camporum var. glandulissima +
Heterotheca camporum +
variety +
110 cm1,100 mm <br />1.1 m <br /> (150 cm1,500 mm <br />1.5 m <br />) +
35 cm350 mm <br />0.35 m <br /> (110 cm1,100 mm <br />1.1 m <br />) +