Heterotheca fulcrata var. amplifolia

(Rydberg) Semple

Rev. Heterotheca Pyllotheca, 74. 1996.

Endemic
Basionyms: Chrysopsis amplifolia Rydberg Bull. Torrey Bot. Club 31: 648. 1905
Synonyms: Chrysopsis foliosa var. amplifolia (Rydberg) A. Nelson Chrysopsis nitidula Wooton & Standley
Treatment appears in FNA Volume 20. Treatment on page 243. Mentioned on page 244, 245, 249.

Stems moderately to densely short-strigose, eglandular or nearly so. Distal cauline leaf-blades ovate to lanceolate, 17–40 × 5–15 mm, margins with few long-hispido-strigose cilia, faces moderately to densely short-strigose (small leaves sometimes nearly white), eglandular or sparsely stipitate-glandular. Subtending bracts ovate, usually large (6–17 × 1.5–5.5 mm), leaflike, well differentiated (sometimes densely strigose when small). Involucres 6.4–10 mm. Phyllaries lanceolate to triangular-lanceolate, faces moderately strigose, usually eglandular. Ray laminae 8.5–14 (–16) mm. Cypsela faces moderately strigose. 2n = 18, 36.


Phenology: Flowering (Jun–)July–Sep(–Oct).
Habitat: Dry gravelly soils, coarse granitic soils, disintegrated granite slopes, high rocky summits, dry grasslands with sagebrush, roadsides, transition zone, Ponderosa pine–oak associations, pinyon-juniper zone at edge of Upper Sonoran Desert, mid to higher montane zone
Elevation: 1300–3500 m

Distribution

V20-533-distribution-map.gif

Ariz., Colo., N.Mex., Utah, Wyo.

Discussion

Variety amplifolia is similar in most respects to var. fulcrata, but the leaf indument is usually densely short-strigose (50–200 hairs/mm2) with few or no glands. Some plants maintain the proximal leaf shape well up the stem, only the few distalmost leaves becoming narrowly ovate-lanceolate; those are similar to Heterotheca villosa var. pedunculata. In southern Utah and eastern Arizona, small-leaved plants may be similar to H. zionensis due to the density of leaf hairs, but in that area the latter typically have densely stipitate-glandular phyllaries, moderately stipitate-glandular distal leaves, and no large bracts subtending the heads.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
corymbiform +, open +  and compact +
indeterminate +  and determinate +
John C. Semple +
(Rydberg) Semple +
rounded +  and attenuate to somewhat +
Chrysopsis amplifolia +
compound +  and simple +
3.4 cm34 mm <br />0.034 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
ovate;lanceolate +
reduced +  and larger +
4mm;15mm +
rugulose +  and muricate +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
long-hispido-strigose +
not 2-lipped +  and actinomorphic +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
1.7 cm17 mm <br />0.017 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
proximal +  and subtending +
Ariz. +, Colo. +, N.Mex. +, Utah +  and Wyo. +
1300–3500 m +
strigose +, stipitate-glandular +  and short-strigose +
differentiated +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Dry gravelly soils, coarse granitic soils,Dry gravelly soils, coarse granitic soils, disintegrated granite slopes, high rocky summits, dry grasslands with sagebrush, roadsides, transition zone, Ponderosa pine–oak associations, pinyon-juniper zone at edge of Upper Sonoran Desert, mid to higher montane zoneSonoran Desert, mid to higher montane zone +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
acute +  and weakly clavate +
tan;rusty brown +
attenuate +  and clavate +
0.9 cm9 mm <br />0.009 m <br /> (1.03 cm10.3 mm <br />0.0103 m <br />) +
cylindric;campanulate +
0.64 cm6.4 mm <br />0.0064 m <br /> (1 cm10 mm <br />0.01 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
1mm;2.7mm +
sessile +  and petiolate +
cauline +  and basal +
erect;spreading +
lanceolate +, triangular +  and deltate +
hispido-strigose +
sinuate +  and entire to somewhat +
2-carpellate +
in 3 +  and 4 series +
stipitate-glandular +  and strigoso-canescent +
0.1 cm1 mm <br />0.001 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering (Jun–)July–Sep(–Oct). +
lanceolate +  and triangular-lanceolate +
ovate-oblanceolate +
subpetiolate +  and sessile +
Rev. Heterotheca Pyllotheca, +
1.4 cm14 mm <br />0.014 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
0.85 cm8.5 mm <br />0.0085 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
10 +  and 12 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
nearly +  and eglandular +
ascending +  and erect +
1 +  and 30 +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis foliosa var. amplifolia +  and Chrysopsis nitidula +
Heterotheca fulcrata var. amplifolia +
Heterotheca fulcrata +
variety +
50 cm500 mm <br />0.5 m <br /> (80 cm800 mm <br />0.8 m <br />) +
17 cm170 mm <br />0.17 m <br /> (50 cm500 mm <br />0.5 m <br />) +
tree +, shrub +  and subshrub +