Heterotheca fulcrata var. fulcrata

Endemic
Synonyms: Chrysopsis cryptocephala Wooton & Standley Chrysopsis elata Osterhout Chrysopsis resinolens A. Nelson Chrysopsis resinolens var. ciliata Chrysopsis viscida subsp. ciliata (A. Nelson) S. F. Blake
Treatment appears in FNA Volume 20. Treatment on page 243. Mentioned on page 244, 254.

Stems sparsely to moderately hispido-strigose and stipitate-glandular. Distal cauline leaf-blades ovate to lanceolate, (12–) 16–39 (–55) × 5–15 mm, margins with few long-hispido-strigose cilia, faces sparsely to moderately hispido-strigose and stipitate-glandular. Subtending bracts ovate, usually large (7–22 × 2–6 mm), leaflike, well differentiated. Involucres 6.7–10.3 mm. Phyllaries lanceolate to triangular-lanceolate, moderately strigose and stipitate-glandular. Ray laminae (8–) 9.8–14.5 mm. Cypsela faces sparsely to moderately strigose. 2n = 18, 36.


Phenology: Flowering (Jun–)Aug–Sep(–Oct).
Habitat: Riparian zones in ponderosa pine and Douglas fir forests, grassy clearings in Ponderosa pine forest, lower spruce-fir zone, sandy riverbanks, roadsides, coarse sandy granitic soils with gravel, crushed lava gravelly soils, moist disturbed soils, shale slopes with sandstone outcrops, open, upper grass and sagebrush slopes
Elevation: 1500–3100 m

Distribution

V20-532-distribution-map.gif

Ariz., Colo., Idaho, Nev., N.Mex., Tex., Utah, Wyo.

Discussion

Variety fulcrata grows in the Front Range of the Rocky Mountains (Colorado and Wyoming), and in disjunct populations in Arizona, New Mexico, and Utah; it is considered very rare and atypical in adjacent Idaho, Nevada, and Texas, where it is of conservation concern. Variety fulcrata is the most variable member of Heterotheca fulcrata. It is characterized by having more hairs than glands per area of leaf face, the hairs being sparse to moderate. Many collections are morphologically intermediate in indument features with var. amplifolia. A few intermediates with the other two varieties also have been collected, as well as a number of possible hybrids with H. pumila and with several varieties of H. villosa.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
corymbiform +, open +  and compact +
indeterminate +  and determinate +
John C. Semple +
(Greene) Shinners +
rounded +  and attenuate to somewhat +
Chrysopsis fulcrata +
compound +  and simple +
3.4 cm34 mm <br />0.034 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
ovate;lanceolate +
reduced +  and larger +
4mm;15mm +
rugulose +  and muricate +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
long-hispido-strigose +
not 2-lipped +  and actinomorphic +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
3.9 cm39 mm <br />0.039 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (3.9 cm39 mm <br />0.039 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
proximal +  and subtending +
Ariz. +, Colo. +, Idaho +, Nev. +, N.Mex. +, Tex. +, Utah +  and Wyo. +
1500–3100 m +
stipitate-glandular +  and hispido-strigose +
differentiated +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Riparian zones in ponderosa pine and DouglRiparian zones in ponderosa pine and Douglas fir forests, grassy clearings in Ponderosa pine forest, lower spruce-fir zone, sandy riverbanks, roadsides, coarse sandy granitic soils with gravel, crushed lava gravelly soils, moist disturbed soils, shale slopes with sandstone outcrops, open, upper grass and sagebrush slopesps, open, upper grass and sagebrush slopes +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
acute +  and weakly clavate +
tan;rusty brown +
attenuate +  and clavate +
0.9 cm9 mm <br />0.009 m <br /> (1.03 cm10.3 mm <br />0.0103 m <br />) +
cylindric;campanulate +
0.67 cm6.7 mm <br />0.0067 m <br /> (1.03 cm10.3 mm <br />0.0103 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
1mm;2.7mm +
sessile +  and petiolate +
cauline +  and basal +
erect;spreading +
lanceolate +, triangular +  and deltate +
hispido-strigose +
sinuate +  and entire to somewhat +
2-carpellate +
in 3 +  and 4 series +
stipitate-glandular +  and strigoso-canescent +
0.1 cm1 mm <br />0.001 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering (Jun–)Aug–Sep(–Oct). +
stipitate-glandular +  and strigose +
lanceolate +  and triangular-lanceolate +
ovate-oblanceolate +
subpetiolate +  and sessile +
Field & Lab. +
0.8 cm8 mm <br />0.008 m <br /> (0.98 cm9.8 mm <br />0.0098 m <br />) +
0.98 cm9.8 mm <br />0.0098 m <br /> (1.45 cm14.5 mm <br />0.0145 m <br />) +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
10 +  and 12 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
ascending +  and erect +
1 +  and 30 +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis cryptocephala +, Chrysopsis elata +, Chrysopsis resinolens +, Chrysopsis resinolens var. ciliata +  and Chrysopsis viscida subsp. ciliata +
Heterotheca fulcrata var. fulcrata +
Heterotheca fulcrata +
variety +
50 cm500 mm <br />0.5 m <br /> (80 cm800 mm <br />0.8 m <br />) +
17 cm170 mm <br />0.17 m <br /> (50 cm500 mm <br />0.5 m <br />) +
tree +, shrub +  and subshrub +