Heterotheca sessiliflora subsp. bolanderi

(A. Gray) Semple

Phytologia 73: 449. 1993.

EndemicConservation concern
Basionym: Chrysopsis bolanderi A. Gray Proc. Amer. Acad. Arts 6: 543. 1866
Synonyms: Chrysopsis arenaria Elmer Chrysopsis villosa var. bolanderi (A. Gray) A. Gray Heterotheca bolanderi (A. Gray) V. L. Harms
Treatment appears in FNA Volume 20. Treatment on page 240. Mentioned on page 238.

Stems sparsely hispido-strigose proximally. Cauline leaves: proximal subsessile to sessile, blades oblanceolate, margins entire, flat or weakly undulate, apices acute, faces usually moderately, rarely densely hispido-strigose, sparsely stipitate-glandular; distal sessile, blades oblanceolate or lanceolate, (13–) 21–40 × (4–) 5–7 (–8) mm, often little reduced distally, margins flat, with large spreading cilia along much of length, faces moderately long-strigose, sparsely to moderately stipitate-glandular. Peduncle bracts 1–4, grading from leaves, lanceolate to linear distally, faces sparsely to moderately long-hispido-strigose, sparsely to moderately stipitate-glandular; sometimes 1–3+ larger leaflike bracts subtending and exceeding involucres. Involucres 9–12.6 (–15) mm. Ray-florets (9–) 12–18 (–21); laminae 9–15 (–20) mm. Disc-florets: corolla-tubes and throats glabrate, lobes glabrate to sparsely strigose (hairs 0.2–0.7 (–1) mm). Cypsela faces moderately to densely strigose.


Phenology: Flowering Jun–Sep(–Oct).
Habitat: Dunes, headlands, open grassy slopes, cliffs, coastal terraces, roadsides in coastal scrub
Elevation: 0–200 m

Discussion

Of conservation concern.

Subspecies bolanderi is known from Mendocino to San Mateo counties and is rare south to n Monterey County near the coast and at lower elevations around San Francisco Bay. The few collections from northwestern Washington came from sandy sites near the ocean. None are typical for subsp. bolanderi, but they would be aberrant in Heterotheca villosa. These were likely chance introductions that did not persist.

Subspecies bolanderi is distinguished by its arrays with few, large heads, flat leaf margins, long hairs, and large cypselae with long pappi bristles. Corolla lobe hairs are often very few and sometimes difficult to find, especially in older heads past flowering. Plants from more upland habitats can be confused with (subsp. echioides) var. bolanderioides, which tends to have easily seen, very long hairs attached to the corolla lobes and tubes. Plants from Mendocino often have long peduncles, while those from around San Fransico Bay tend to have shorter peduncles and more congested arrays.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
paniculiform +  and corymbiform +
indeterminate +  and determinate +
John C. Semple +
(A. Gray) Semple +
rounded +  and cuneate +
Chrysopsis bolanderi +
compound +  and simple +
1.3 cm13 mm <br />0.013 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
2.1 cm21 mm <br />0.021 m <br /> (4 cm40 mm <br />0.04 m <br />) +
lanceolate;oblanceolate;oblanceolate +
reduced +
5mm;7mm +
rugulose +  and muricate +
leafy +  and phyllary-like +
0.85 cm8.5 mm <br />0.0085 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
barbellulate +  and barbellate +
0.8 cm8 mm <br />0.008 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.7 cm7 mm <br />0.007 m <br /> (0.95 cm9.5 mm <br />0.0095 m <br />) +
glabrous +  and glabrate +
ampliate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
0–200 m +
long-hispido-strigose +  and sparsely moderately stipitate-glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Dunes, headlands, open grassy slopes, cliffs, coastal terraces, roadsides in coastal scrub +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
lanceolate +
reduced +
each +  and sessile +
attenuate +  and weakly clavate +
tan;rusty brown +
attenuate +  and clavate +
1.26 cm12.6 mm <br />0.0126 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
campanulate;turbinate;campanulate;turbinate;cylindric +
0.9 cm9 mm <br />0.009 m <br /> (1.26 cm12.6 mm <br />0.0126 m <br />) +
1.05 cm10.5 mm <br />0.0105 m <br /> (1.85 cm18.5 mm <br />0.0185 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.24 cm2.4 mm <br />0.0024 m <br />) +
4.5mm;10.5mm +
0.9 cm9 mm <br />0.009 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1mm;1.7mm +
sessile +  and petiolate +
proximal +  and basal +
erect;spreading +
glabrate +  and sparsely strigose +
lanceolate +, triangular +  and deltate +
fimbriate-ciliate +  and strigoso-ciliate +
stipitate-glandular +  and strigose +
undulate +, flat +  and entire +
2-carpellate +
in 3 +  and 4 series +
glandular +, strigose +  and hispid +
0.2 cm2 mm <br />0.002 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Flowering Jun–Sep(–Oct). +
lanceolate +  and linear-lanceolate +
ovate-oblanceolate +
subsessile +  and sessile +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
6 +  and 10 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
exalbuminous +
Endemic +  and Conservation concern +
decumbent +  and ascending or erect +
1 +  and 40 +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis arenaria +, Chrysopsis villosa var. bolanderi +  and Heterotheca bolanderi +
Heterotheca sessiliflora subsp. bolanderi +
Heterotheca sessiliflora +
subspecies +
70 cm700 mm <br />0.7 m <br /> (110 cm1,100 mm <br />1.1 m <br />) +
20 cm200 mm <br />0.2 m <br /> (70 cm700 mm <br />0.7 m <br />) +
tree +, shrub +  and subshrub +