Heterotheca stenophylla var. angustifolia

(Rydberg) Semple

Novon 4: 53. 1994.

Endemic
Basionym: Chrysopsis angustifolia Rydberg Bull. Torrey Bot. Club 37: 128. 1910
Synonyms: Chrysopsis villosa var. angustifolia (Rydberg) Cronquist Heterotheca villosa var. angustifolia (Rydberg) V. L. Harms
Treatment appears in FNA Volume 20. Treatment on page 247. Mentioned on page 246, 248, 249.

Distal mid-stems moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular. Leaves pale green to gray-green, faces moderately to ± densely hispido-strigose (21–60 hairs/mm²; hairs rarely broad-based, pustulate), eglandular to sparsely glandular; distal cauline linear to narrowly oblanceolate, rarely broadly so, (14–) 18.7 (–30) × (2–) 2.6–4.5 (–6) mm, margins proximally long-hispido-strigose, sometimes cilia numerous along whole length, faces sparsely to moderately hispido-strigose, eglandular. Phyllary faces usually moderately strigose, eglandular. 2n = 36.


Phenology: Flowering May–Aug(–Oct).
Habitat: Sandy and gravelly soils, sandy loam, sandy gypsiferous soil, prairies, hillsides, pastures, open areas by oak wooded ravines, disturbed slopes, grasslands, stabilized dunes
Elevation: 300–1400 m

Distribution

V20-541-distribution-map.gif

Colo., Kans., Nebr., N.Mex., Okla., S.Dak., Tex.

Discussion

Variety angustifolia grows in the Great Plains from central Texas and New Mexico to eastern Colorado and southern South Dakota. Triploid chromosome counts from putative intervarietal hybrids have been reported. Variety angustifolia usually has densely hispid distal mid stems with many long-spreading hairs and densely hispido-strigose, eglandular leaves and phyllaries. The most densely hairy individuals are not always easily distinguished from sympatric forms of Heterotheca canescens. Both diploid and tetraploid individuals of H. canescens usually have more than 100 hairs/mm2 of leaf face versus less than 60 hairs/mm2 in both varieties of H. stenophylla. The difference manifests itself in silvery gray-green versus green (bright to dark) or gray-green leaves, respectively. Stems of H. villosa usually have relatively longer nodes and a less densely leafy appearance. The leaves can be as densely hairy. Variety angustifolia historically has been included in H. villosa.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
John C. Semple +
(Rydberg) Semple +
cuneate +  and attenuate +
Chrysopsis angustifolia +
compound +  and simple +
0.9 cm9 mm <br />0.009 m <br /> (1.65 cm16.5 mm <br />0.0165 m <br />) +
0.43 cm4.3 mm <br />0.0043 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
1.65 cm16.5 mm <br />0.0165 m <br /> (3 cm30 mm <br />0.03 m <br />) +
linear-oblanceolate;narrowly oblanceolate +
reduced +
2.2mm;4.3mm +
rugulose +  and muricate +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
additional +
not 2-lipped +  and actinomorphic +
ampliate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
1.87 cm18.7 mm <br />0.0187 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
linear +  and narrowly oblanceolate +
0.26 cm2.6 mm <br />0.0026 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Colo. +, Kans. +, Nebr. +, N.Mex. +, Okla. +, S.Dak. +  and Tex. +
300–1400 m +
eglandular +  and sparsely glandular +
hispido-strigose +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Sandy and gravelly soils, sandy loam, sandy gypsiferous soil, prairies, hillsides, pastures, open areas by oak wooded ravines, disturbed slopes, grasslands, stabilized dunes +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.8 cm8 mm <br />0.008 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
cylindro-turbinate;narrowly campanulate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
1.15 cm11.5 mm <br />0.0115 m <br /> (1.65 cm16.5 mm <br />0.0165 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
7mm;11.5mm +
0.7mm;1.5mm +
sessile +  and petiolate +
pale green +  and gray-green +
cauline +  and basal +
erect;spreading +
lanceolate +, triangular +  and deltate +
long-hispido-strigose +
2-carpellate +
in 3 +  and 4 series +
stipitate-glandular +  and hispido-strigose +
0.2 cm2 mm <br />0.002 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
Flowering May–Aug(–Oct). +
narrowly triangular-lanceolate +  and lanceolate +
ovate-oblanceolate +
linear-oblanceolate +  and oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
6 +  and 10 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
exalbuminous +
short-branched +
ascending +  and erect +
1 +  and 50 +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis villosa var. angustifolia +  and Heterotheca villosa var. angustifolia +
Heterotheca stenophylla var. angustifolia +
Heterotheca stenophylla +
variety +
46 cm460 mm <br />0.46 m <br /> (65 cm650 mm <br />0.65 m <br />) +
18 cm180 mm <br />0.18 m <br /> (46 cm460 mm <br />0.46 m <br />) +
tree +, shrub +  and subshrub +