Heterotheca villosa var. ballardii

(Rydberg) Semple

Novon 4: 53. 1994.

Endemic
Basionyms: Chrysopsis ballardii Rydberg Brittonia 1: 100. 1931
Treatment appears in FNA Volume 20. Treatment on page 251. Mentioned on page 250.

Stems ascending to erect, 22–50 cm, moderately to densely strigose, eglandular, often abundantly long-hirsute. Distal cauline leaf-blades oblong, (18–) 22–34 (–42) × 4.3–9.3 (–12.5) mm, little reduced distally, bases rounded, margins flat, faces moderately strigose (hairs usually not obscuring faces, 20–56 (–70) /mm2), eglandular. Heads (2–) 4–16 (–24), in corymbiform arrays. Peduncles (5–) 12–47 (–67) mm, moderately to densely hispido-strigose, eglandular; bracts subtending heads leafy, oblanceolate, often exceeding involucres. Involucres broadly campanulate, (5.5–) 6.5–9.3 mm. Phyllaries narrowly triangular-lanceolate, moderately to densely strigose, eglandular to sparsely glandular, margins often reddish purple distally. Ray-florets (12–) 17–30 (avereaging 21), laminae (8.5–) 9.9–14.5 mm (averaging 11.5 mm; showy). 2n = 36.


Phenology: Flowering (Jun–)Jul–Aug(–Oct).
Habitat: Dry sandy clay, thin silty loam, sandy loam, gravel soils, dry, rocky, granite-gravel soils, prairies, fields, roadsides, railroad rights-of-way
Elevation: 300–2300 m

Distribution

V20-546-distribution-map.gif

Alta., Man., Ont., Sask., Colo., Minn., Mont., Nebr., N.Dak., S.Dak.

Discussion

Variety ballardii grows in the northeastern Great Plains; it is rare and probably introduced in Colorado, Montana, and northwestern Ontario. In its typical, large-leaved and many-rayed form, var. ballardii is quite distinct. It is distinguished by its rounded leaf bases, indument of short hairs, eglandular, and large, many-rayed heads often subtended by one or more large, narrow, leaflike peduncle bracts. Plants of var. ballardii are sufficiently similar to those of var. foliosa that the two taxa might be treated as convarietal under the name var. foliosa. V. L. Harms (2003) treated the Saskatchewan plants as members of a more broadly defined var. villosa.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
John C. Semple +
(Rydberg) Semple +
rounded +
Chrysopsis ballardii +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
3.5mm;7mm +
rugulose +  and muricate +
oblanceolate +
reduced +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
3.4 cm34 mm <br />0.034 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.93 cm9.3 mm <br />0.0093 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
2.2 cm22 mm <br />0.022 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
0.43 cm4.3 mm <br />0.0043 m <br /> (0.93 cm9.3 mm <br />0.0093 m <br />) +
Alta. +, Man. +, Ont. +, Sask. +, Colo. +, Minn. +, Mont. +, Nebr. +, N.Dak. +  and S.Dak. +
300–2300 m +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Dry sandy clay, thin silty loam, sandy loam, gravel soils, dry, rocky, granite-gravel soils, prairies, fields, roadsides, railroad rights-of-way +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
campanulate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.93 cm9.3 mm <br />0.0093 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.85 cm8.5 mm <br />0.0085 m <br /> (0.99 cm9.9 mm <br />0.0099 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.99 cm9.9 mm <br />0.0099 m <br /> (1.45 cm14.5 mm <br />0.0145 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
4.7 cm47 mm <br />0.047 m <br /> (6.7 cm67 mm <br />0.067 m <br />) +
hispido-strigose +
1.2 cm12 mm <br />0.012 m <br /> (4.7 cm47 mm <br />0.047 m <br />) +
Flowering (Jun–)Jul–Aug(–Oct). +
eglandular +  and sparsely glandular +
triangular-lanceolate +
ovate-oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
ascending +  and erect +
1 +  and 50 +
22 cm220 mm <br />0.22 m <br /> (50 cm500 mm <br />0.5 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis villosa +, Diplogon villosum +  and Diplopappus villosus +
Heterotheca villosa var. ballardii +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +