Heterotheca villosa var. depressa

(Rydberg) Semple

Novon 4: 53. 1994.

EndemicConservation concern
Basionyms: Chrysopsis depressa Rydberg Mem. New York Bot. Gard. 1: 381. 1900
Synonyms: Heterotheca depressa (Rydberg) Dorn
Treatment appears in FNA Volume 20. Treatment on page 251. Mentioned on page 248, 250, 252.

Stems ascending to erect, 5–22 cm, moderately to densely hispido-strigose, sparsely long-hirsute, eglandular or sparsely stipitate-glandular. Distal cauline leaf-blades linear-oblanceolate, 5–19 (–23) × 2.3–4.2 (–4.7) mm, sometimes little reduced, bases cuneate to attenuate, margins flat, apices acute, faces densely strigose (hairs 84–140/mm2), sparsely to to moderately stipitate-glandular (glands 0–28/mm2). Heads 1–8 (–10) in congested to open corymbiform arrays. Peduncles (7–) 12–28 (–37) mm, moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular; bracts subtending heads small or none. Involucres cylindric to narrowly campanulate (fresh), 5.5–6.8 (–7.5) mm. Phyllaries lanceolate to narrowly triangular-lanceolate, margins often reddish purple distally, faces moderately to densely strigose, eglandular or sparsely stipitate-glandular. Ray-florets 5–13 (–15); laminae 5–9.5 (–11) mm. 2n = 36.


Phenology: Flowering (Jun–)Jul–Sep(–Nov).
Habitat: Hot springs, broken travertine on old terraces, dry geyserite, gravelly stream banks, dry rocky hillsides, open slopes, dry sandy soils, montane conifer forests
Elevation: 1800–2400 m

Discussion

Of conservation concern.

Variety depressa grows in geyser basins in Yellowstone National Park and is locally common in rocky soils in Teton National Park; it is very rare farther south in Wyoming. It is distinguished by its small stature, small heads, and linear-oblanceolate leaves with many hairs and usually some glands. It is similar to some forms of var. minor, which occur throughout the range of var. depressa and with which it hybridizes. It is also similar to hairier forms of var. villosa. Typical var. depressa occupies the vicinity of hot springs and geyser basins and nearby river flood banks, while var. minor occupies other drier or non-geyserite nearby habitats.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
corymbiform +, congested +  and open +
indeterminate +  and determinate +
John C. Semple +
(Rydberg) Semple +
cuneate +  and attenuate +
Chrysopsis depressa +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
3.5mm;7mm +
rugulose +  and muricate +
oblanceolate +  and linear-oblanceolate +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
1.9 cm19 mm <br />0.019 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
0.42 cm4.2 mm <br />0.0042 m <br /> (0.47 cm4.7 mm <br />0.0047 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.9 cm19 mm <br />0.019 m <br />) +
0.23 cm2.3 mm <br />0.0023 m <br /> (0.42 cm4.2 mm <br />0.0042 m <br />) +
1800–2400 m +
stipitate-glandular +  and strigose +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Hot springs, broken travertine on old terraces, dry geyserite, gravelly stream banks, dry rocky hillsides, open slopes, dry sandy soils, montane conifer forests +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.68 cm6.8 mm <br />0.0068 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
cylindric;narrowly campanulate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.68 cm6.8 mm <br />0.0068 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.95 cm9.5 mm <br />0.0095 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.5 cm5 mm <br />0.005 m <br /> (0.95 cm9.5 mm <br />0.0095 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
2.8 cm28 mm <br />0.028 m <br /> (3.7 cm37 mm <br />0.037 m <br />) +
stipitate-glandular +  and hispido-strigose +
1.2 cm12 mm <br />0.012 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
Flowering (Jun–)Jul–Sep(–Nov). +
lanceolate +  and narrowly triangular-lanceolate +
ovate-oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
Endemic +  and Conservation concern +
ascending +  and erect +
1 +  and 50 +
5 cm50 mm <br />0.05 m <br /> (22 cm220 mm <br />0.22 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Heterotheca depressa +
Heterotheca villosa var. depressa +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +