Heterotheca villosa var. foliosa

(Nuttall) V. L. Harms

Wrightia 4: 15. 1968.

Endemic
Basionyms: Chrysopsis foliosa Nuttall Trans. Amer. Philos. Soc., n. s. 7: 316. 1840
Synonyms: Chrysopsis butleri Rydberg Chrysopsis foliosa var. imbricata (A. Nelson) A. Nelson Chrysopsis imbricata (Pursh) Nuttall ex de Candolle Chrysopsis villosa var. foliosa (Nuttall) Cronquist Heterotheca foliosa
Treatment appears in FNA Volume 20. Treatment on page 251. Mentioned on page 249, 250, 254.

Stems ascending to erect, 22–41 (–58) cm, moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular, often abundantly long-hirsute. Distal cauline leaf-blades oblong, rarely ovate, 11–28 (–33) × 3.5–8.5 (–9.7) mm, sometimes little reduced distally, bases rounded, margins flat, apices broadly acute to obtuse, faces moderately to densely hispido-strigose, (hairs sometimes obscuring faces, 25–59 (–90) mm2), eglandular or sparsely stipitate-glandular. Heads 4–24 (–35) in congested to open, corymbiforms arrays. Peduncles (9–) 17–39 mm, moderately to densely hispido-strigose, eglandular; bracts subtending heads sometimes oblanceolate, leafy, often surpassing them. Involucres narrowly campanulate (fresh), 6–7.6 (–8.6) mm. Phyllaries narrowly triangular-lanceolate, margins often reddish purple distally, faces moderately to densely strigose, eglandular to very sparsely glandular. Ray-florets (7–) 13–22, laminae 8–11.4 (–12.4) mm (less showy). 2n = 18, 36.


Phenology: Flowering Jul–Sep(–Oct).
Habitat: Sandy shale and clay soils, sandy loam on sandstone, gypsum soils, crevices in granite and sandstone, loose talus slopes, stabilized dunes, dry coulees and flood plains, dry short-grass and semibarren prairies, grassy areas with scattered sagebrush, Ponderosa pine, aspen, and Cercocarpus, roadsides
Elevation: 600–2600 m

Distribution

V20-547-distribution-map.gif

Alta., Sask., Colo., Idaho, Mont., Nebr., N.Mex., N.Dak., Oreg., S.Dak., Wash.

Discussion

Variety foliosa is most common at the base of the Front Range in Colorado and Wyoming; it occurs scattered across the northern part of its range in the mountains and western prairies. Plants that are glandular but otherwise like var. foliosa are presumed here to be hybrids with either var. minor or var. nana. Variety foliosa is most similar to var. ballardii. V. L. Harms (2003) treated Saskatchwan plants as members of more broadly defined var. villosa.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
congested +  and open +
indeterminate +  and determinate +
corymbiforms +
John C. Semple +
(Nuttall) V. L. Harms +
rounded +
Chrysopsis foliosa +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
3.5mm;7mm +
rugulose +  and muricate +
oblanceolate +
reduced +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
2.8 cm28 mm <br />0.028 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
0.85 cm8.5 mm <br />0.0085 m <br /> (0.97 cm9.7 mm <br />0.0097 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
Alta. +, Sask. +, Colo. +, Idaho +, Mont. +, Nebr. +, N.Mex. +, N.Dak. +, Oreg. +, S.Dak. +  and Wash. +
600–2600 m +
eglandular to very +
strigose +  and hispido-strigose +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Sandy shale and clay soils, sandy loam on Sandy shale and clay soils, sandy loam on sandstone, gypsum soils, crevices in granite and sandstone, loose talus slopes, stabilized dunes, dry coulees and flood plains, dry short-grass and semibarren prairies, grassy areas with scattered sagebrush, Ponderosa pine, aspen, and Cercocarpus, roadsidessa pine, aspen, and Cercocarpus, roadsides +
stipitate-glandular +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.76 cm7.6 mm <br />0.0076 m <br /> (0.86 cm8.6 mm <br />0.0086 m <br />) +
campanulate +
0.6 cm6 mm <br />0.006 m <br /> (0.76 cm7.6 mm <br />0.0076 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.14 cm11.4 mm <br />0.0114 m <br /> (1.24 cm12.4 mm <br />0.0124 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.8 cm8 mm <br />0.008 m <br /> (1.14 cm11.4 mm <br />0.0114 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
0.9 cm9 mm <br />0.009 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
hispido-strigose +
1.7 cm17 mm <br />0.017 m <br /> (3.9 cm39 mm <br />0.039 m <br />) +
Flowering Jul–Sep(–Oct). +
triangular-lanceolate +
ovate-oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
ascending +  and erect +
1 +  and 50 +
22 cm220 mm <br />0.22 m <br /> (41 cm410 mm <br />0.41 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis butleri +, Chrysopsis foliosa var. imbricata +, Chrysopsis imbricata +, Chrysopsis villosa var. foliosa +  and Heterotheca foliosa +
Heterotheca villosa var. foliosa +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +