Heterotheca villosa var. minor

(Hooker) Semple

Novon 4: 54. 1994.

IllustratedEndemic
Basionym: Chrysopsis villosa var. minor Hooker London J. Bot. 6: 244. 1847
Synonyms: Chrysopsis arida A. Nelson Chrysopsis asprella Greene Chrysopsis bakeri Greene Chrysopsis columbiana Greene Chrysopsis compacta Greene Chrysopsis floribunda Greene Chrysopsis grandis Rydberg Chrysopsis hirsuta Greene Chrysopsis hirsutissima Greene Chrysopsis hispida (Hooker) de Candolle Chrysopsis villosa var. discoidea A. Gray Chrysopsis villosa var. hispida (Hooker) A. Gray Chrysopsis wisconsinensis (Pursh) Kuntze Diplogon villosum var. discoideum (A. Gray) Kuntze Diplopappus hispidus (Pursh) Shinners Heterotheca villosa var. hispida (Hooker) V. L. Harms
Treatment appears in FNA Volume 20. Treatment on page 252. Mentioned on page 244, 245, 249, 251, 253, 25.

Stems decumbent to ascending-erect, (9–) 16–33 (–48) mm, sparsely to densely hispido-strigose, sparsely to abundantly long-hirsute, sparsely to densely stipitate-glandular. Distal cauline leaf-blades usually narrowly to broadly oblanceolate or oblong, sometimes ovate (in mountains), (11–) 16–28 (–40) mm, ± reduced distally, bases usually narrowly to broadly convex-cuneate to attenuate, sometimes rounded, margins flat, sometimes remotely undulate, apices obtuse or acute, faces sparsely to moderately hispido-strigose, (1–75/mm2), sparsely to moderately glandular (glands 1–32/mm2). Heads 1–13 (–42) in congested to open, corymbiform arrays, branches usually not very long. Peduncles 4.5–40 (–60) mm, moderately to densely hispido-strigose, usually sparsely to moderately stipitate-glandular rarely eglandular; bracts usually linear-oblong, reduced, rarely linear-lanceolate and leaflike, sparsely to moderately hispido-strigose, sparsely to moderately stipitate-glandular. Involucres cylindric to campanulate (fresh), (5.5–) 6.5–8.5 (–9.3) mm. Phyllaries narrowly triangular-lanceolate, faces usually sparsely, rarely densely strigose (hairs to 1 mm), usually sparsely to moderately stipitate-glandular, rarely eglandular. Ray-florets (7–) 10–18 (–26), laminae (4–) 6–10 (–12.6) mm. 2n = 18, 36.


Phenology: Flowering (May–)Jun–Aug(–Nov).
Habitat: Gravelly, sandy, and loamy soils, crevices in granite, limestone rocks, marble rocks, eroded granites and sandstones, basaltic cliffs, lava flows, rocky slopes, dry ledges, stream banks, glacial outwashes, roadside railroad embankments, grasslands, ponderosa pine-oak woods, open pinyon-juniper associations
Elevation: (60–)300–2900(–3200) m

Distribution

V20-550-distribution-map.gif

Alta., B.C., Man., Sask., Ariz., Calif., Colo., Idaho., Ill., Iowa, Kans., Minn., Mont., Nebr., Nev., N.Mex., N.Dak., Oreg., S.Dak., Utah, Wash., Wis., Wyo.

Discussion

Variety minor is distinguished by its usually narrowly to broadly oblancelate distal cauline leaves (sometime oblong or ovate) and moderately glandular and hispido-strigose indument. The leaf bases are usually narrowly to broadly cuneate or attenuate (rarely rounded). Distal leaf faces usually have about 10–50 hairs/mm2 and about 4–20 glands/mm2. The variety is the most variable in the species and includes some local distinctive morphotypes that grade into other forms. Plants intermediate between this and all other varieties occur in areas where the ranges are sympatric, and they make infraspecific taxonomy of the species difficult. The variety has been incorrectly referred to as var. hispida (a later synonym) in most floras.

The status of Heterotheca barbata (Rydberg) Semple (Chrysopsis barbata Rydberg), the Spokane goldenaster, is uncertain. J. C. Semple (1996) treated it as a separate species to draw attention to the problem; a detailed description based on the type and detailed illustrations were included. It is known from the type collection along the Spokane River Valley east of Spokane, Idaho, and two down-river, atypical collections (Benton and Spokane counties, Washington). It flowers in July (sometimes August). It is similar to H. villosa var. minor, but differs in having lanceolate-elliptic distal cauline leaves (34–38 × 8–9 mm) that are little reduced distally, long branches (each with one to a few large heads), and disc corollas with a few, very long hairs on the tube. Further work is needed to increase the number of specimens available for a detailed comparison with H. villosa var. minor in order to clarify whether H. barbata warrants species level recognition, should be included in H. villosa as a variety, or placed in synonymy under var. minor.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
acute;obtuse +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
corymbiform +, congested +  and open +
indeterminate +  and determinate +
John C. Semple +
(Hooker) Semple +
rounded +, convex-cuneate +  and attenuate +
Chrysopsis villosa var. minor +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
rugulose +  and muricate +
stipitate-glandular +  and hispido-strigose +
linear-lanceolate +  and linear-oblong +
reduced +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
2.8 cm28 mm <br />0.028 m <br /> (4 cm40 mm <br />0.04 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
ovate +, oblong +  and oblanceolate +
Alta. +, B.C. +, Man. +, Sask. +, Ariz. +, Calif. +, Colo. +, Idaho. +, Ill. +, Iowa +, Kans. +, Minn. +, Mont. +, Nebr. +, Nev. +, N.Mex. +, N.Dak. +, Oreg. +, S.Dak. +, Utah +, Wash. +, Wis. +  and Wyo. +
(60–)300–2900(–3200) m +
stipitate-glandular +, strigose +, punct +  and sparsely moderately glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Gravelly, sandy, and loamy soils, crevicesGravelly, sandy, and loamy soils, crevices in granite, limestone rocks, marble rocks, eroded granites and sandstones, basaltic cliffs, lava flows, rocky slopes, dry ledges, stream banks, glacial outwashes, roadside railroad embankments, grasslands, ponderosa pine-oak woods, open pinyon-juniper associationsak woods, open pinyon-juniper associations +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.85 cm8.5 mm <br />0.0085 m <br /> (0.93 cm9.3 mm <br />0.0093 m <br />) +
cylindric;campanulate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.26 cm12.6 mm <br />0.0126 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
stipitate-glandular +  and hispido-strigose +
0.45 cm4.5 mm <br />0.0045 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Flowering (May–)Jun–Aug(–Nov). +
triangular-lanceolate +
ovate-oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
Illustrated +  and Endemic +
3.3 cm33 mm <br />0.033 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
decumbent +  and ascending-erect +
hispido-strigose +  and sparsely abundantly long-hirsute +
1 +  and 50 +
1.6 cm16 mm <br />0.016 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis arida +, Chrysopsis asprella +, Chrysopsis bakeri +, Chrysopsis columbiana +, Chrysopsis compacta +, Chrysopsis floribunda +, Chrysopsis grandis +, Chrysopsis hirsuta +, Chrysopsis hirsutissima +, Chrysopsis hispida +, Chrysopsis villosa var. discoidea +, Chrysopsis villosa var. hispida +, Chrysopsis wisconsinensis +, Diplogon villosum var. discoideum +, Diplopappus hispidus +  and Heterotheca villosa var. hispida +
Heterotheca villosa var. minor +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +