Heterotheca villosa var. pedunculata

(Greene) V. L. Harms ex Semple

Brittonia 39: 383. 1987.

Endemic
Basionyms: Chrysopsis pedunculata Greene Pittonia 4: 154. 1900
Treatment appears in FNA Volume 20. Treatment on page 252. Mentioned on page 234, 244, 249.

Stems erect, (12–) 30–50 (–75) cm, moderately to densely strigose, usually sparsely, rarely abundantly long-hirsute eglandular or sparsely stipitate-glandular. Distal cauline leaf-blades oblanceolate, (4–) 14–31 × (1.5) –3.7–8 mm, sometimes little reduced distally, margins flat, faces (pale green to white) densely strigose (hairs (42–270) /mm2), eglandular or sparsely stipitate-glandular (glands 20–49/mm2). Heads 1–18 (–39), in congested to open, usually corymbiform, sometimes subumbelliform arrays. Peduncles (7–) 12–40 (–58) mm, moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular; bracts few, linear-oblong, reduced distally, bracts subtending heads phyllary-like or none. Involucres cylindric to narrowly campanulate (fresh), (4.6–) 6.6–11 mm. Phyllaries lanceolate to narrowly triangular-lanceolate, margins often reddish purple distally, faces densely strigose, eglandular to sparsely glandular. Ray-florets 9–17 (–26), laminae (5.5–) 6.6–10.5 (–12) mm. 2n = 18, 36.


Phenology: Flowering (May–)Jun–Oct(–Nov in s).
Habitat: Sandy soils, ponderosa and fir-spruce forests, Upper Sonoran Zone (pinyon-juniper), mixed grasslands, disturbed areas, rocky outcrops, pastures, riparian habitats
Elevation: 1200–2900 m

Distribution

V20-549-distribution-map.gif

Ariz., Colo., N.Mex., Utah.

Discussion

Variety pedunculata usually has pale green to white, densely strigose, sparsely to moderately stipitate-glandular, oblanceolate leaves, and often long-pedunculate heads that are not subtended by large bracts. It hybridizes with var. minor throughout its range, and is similar to but more densely hairy than the more northern and eastern var. villosa.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
paniculiform +, corymbiform +  and open +
indeterminate +  and determinate +
John C. Semple +
(Greene) V. L. Harms ex Semple +
attenuate +  and convex-cuneate +
Chrysopsis pedunculata +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
3.5mm;7mm +
rugulose +  and muricate +
phyllary-like +
linear-oblong +
reduced +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
Ariz. +, Colo. +, N.Mex. +  and Utah. +
1200–2900 m +
eglandular +  and sparsely glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Sandy soils, ponderosa and fir-spruce forests, Upper Sonoran Zone (pinyon-juniper), mixed grasslands, disturbed areas, rocky outcrops, pastures, riparian habitats +
stipitate-glandular +
subumbelliform +, corymbiform +, congested +  and open +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.46 cm4.6 mm <br />0.0046 m <br /> (0.66 cm6.6 mm <br />0.0066 m <br />) +
cylindric;narrowly campanulate +
0.66 cm6.6 mm <br />0.0066 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.05 cm10.5 mm <br />0.0105 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.66 cm6.6 mm <br />0.0066 m <br /> (1.05 cm10.5 mm <br />0.0105 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
4 cm40 mm <br />0.04 m <br /> (5.8 cm58 mm <br />0.058 m <br />) +
stipitate-glandular +  and hispido-strigose +
1.2 cm12 mm <br />0.012 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Flowering (May–)Jun–Oct(–Nov in s). +
lanceolate +  and narrowly triangular-lanceolate +
ovate-oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
1 +  and 50 +
30 cm300 mm <br />0.3 m <br /> (50 cm500 mm <br />0.5 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis villosa +, Diplogon villosum +  and Diplopappus villosus +
Heterotheca villosa var. pedunculata +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +