Heterotheca villosa var. scabra

(Eastwood) Semple

Phytologia 73: 453. 1993.

Endemic
Basionyms: Chrysopsis villosa var. scabra Eastwood Proc. Calif. Acad. Sci., ser. 2, 6: 294. 1896
Synonyms: Chrysopsis viscida subsp. cinerascens S. F. Blake Heterotheca horrida subsp. cinerascens (S. F. Blake) Semple
Treatment appears in FNA Volume 20. Treatment on page 253. Mentioned on page 234, 244, 248, 250, 254, 255.

Stems ascending to erect, 20–44 (–52) cm, moderately to densely hispido-strigose, abundantly long-hirsute, moderately to densely stipitate-glandular. Distal cauline leaf (often perpendicular to stems) blades usually lanceolate to triangular-lanceolate, rarely narrowly ovate, 12–34 × 3–8 mm, somewhat to much reduced distally, margins flat, apices usually sharply acute, faces sparsely to moderately densely hispido-strigose (hairs 5–57, sometimes to 90/mm2), sparsely to densely stipitate-glandular (glands 8–50 mm2). Heads 1–19 (–35), borne singly or in usually very open, sometimes compact, corymbiform arrays, branches often very long. Peduncles (14–) 24–47 (–120) mm, moderately to densely hispido-strigose, sparsely to densely stipitate-glandular; bracts grading from distal leaves, reduced distally, rarely leaflike. Involucres narrowly campanulate (fresh), (6–) 6.8–9.7 (–11) mm. Phyllaries narrowly triangular-lanceolate, margins rarely reddish purple distally, faces sparsely to moderately short-strigose, usually moderately to densely, rarely sparsely stipitate-glandular. Ray-florets (7–) 9–15 (–18), laminae 5–10.6 (12.4) mm. 2n = 18, 36.


Phenology: Flowering (Jan, May–)Jun–Oct(–Nov, south).
Habitat: Lava crevices, limestone cliffs, rocky sandstone loams, red sandstone soils, rocky embankments, rocky slopes, rock outcrops, dry rocky washes, sand flats near streams, in oak woods, juniper-pinyon scrub
Elevation: 400–3200 m

Distribution

V20-551-distribution-map.gif

Ariz., Calif., Idaho, Nev., Utah.

Discussion

Variety scabra is disjunct in the Little San Bernardino Mountains (California). It has moderately to densely stipitate-glandular and sparsely to moderately hispido-strigose, narrowly triangular-lanceolate leaves, and heads with very sparsely hispido-strigose, moderately to densely stipitate-glandular phyllaries. The variation in leaf traits is related to differences in growing conditions and time of the year shoots are produced. Plants may bloom early in the year and again in the late summer or fall. Early blooming plants can be quite different in array form and leaf indument traits. In southwestern Utah and north-central Arizona, var. scabra and var. nana can be difficult to distinguish. In much of Utah, northern Nevada, and adjacent Idaho, var. scabra and var. minor can be difficult to distinguish.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
paniculiform +, corymbiform +  and open +
indeterminate +  and determinate +
John C. Semple +
(Eastwood) Semple +
attenuate +  and convex-cuneate +
Chrysopsis villosa var. scabra +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
3.5mm;7mm +
rugulose +  and muricate +
oblanceolate +  and linear-oblanceolate +
reduced +
corymbiform +  and open +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
Ariz. +, Calif. +, Idaho +, Nev. +  and Utah. +
400–3200 m +
stipitate-glandular +, short-strigose +, moderately densely hispido-strigose +  and sparsely densely stipitate-glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Lava crevices, limestone cliffs, rocky sandstone loams, red sandstone soils, rocky embankments, rocky slopes, rock outcrops, dry rocky washes, sand flats near streams, in oak woods, juniper-pinyon scrub +
paniculiform +
in usually very open , sometimes compact , corymbiform arrays , branches +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.97 cm9.7 mm <br />0.0097 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
campanulate +
0.68 cm6.8 mm <br />0.0068 m <br /> (0.97 cm9.7 mm <br />0.0097 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
12.4 cm124 mm <br />0.124 m <br /> (?) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.5 cm5 mm <br />0.005 m <br /> (1.06 cm10.6 mm <br />0.0106 m <br />) +
sessile +  and petiolate +
1.2 cm12 mm <br />0.012 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
4.7 cm47 mm <br />0.047 m <br /> (12 cm120 mm <br />0.12 m <br />) +
stipitate-glandular +  and hispido-strigose +
2.4 cm24 mm <br />0.024 m <br /> (4.7 cm47 mm <br />0.047 m <br />) +
Flowering (Jan, May–)Jun–Oct(–Nov, south). +
triangular-lanceolate +
ovate-oblanceolate +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
ascending +  and erect +
1 +  and 50 +
20 cm200 mm <br />0.2 m <br /> (44 cm440 mm <br />0.44 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis viscida subsp. cinerascens +  and Heterotheca horrida subsp. cinerascens +
Heterotheca villosa var. scabra +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +