Heterotheca villosa var. villosa

Endemic
Synonyms: Chrysopsis mollis Nuttall
Treatment appears in FNA Volume 20. Treatment on page 250. Mentioned on page 249, 251, 252, 256.

Stems decumbent to erect, 16–39 cm, moderately to densely strigose, sparsely to abundantly long-hirsute, eglandular or sparsely stipitate-glandular. Distal cauline leaf-blades oblanceolate, 21–35 × 3.5–6 mm, bases narrowly cuneate to attenuate, margins flat, apices acute, faces moderately strigose (20–60 hairs/mm2), eglandular or sparsely stipitate-glandular. Heads 1–12 (–25). Peduncles (5–) 12–47 (–67) mm, sparsely to densely hispido-strigose, eglandular; bracts subtending heads usually none, rarely linear-oblanceolate, leaflike. Involucres narrowly cylindric to campanulate (fresh), (5–) 6–9 (–11) mm. Phyllaries narrowly triangular-lanceolate, margins usually reddish purple distally, faces moderately to densely strigose, eglandular. Ray-florets 10–27 (–38), laminae (6.5–) 8.5–15 (–20) mm. 2n = 18, 36.


Phenology: Flowering (May–)Jun–Oct(–Nov, south).
Habitat: Silty, sandy loam or clay soils, chalk, or granitic soils, gravel soils, dry shale-limestone soils, red sandstone soils, travertine soils of sandhill prairies, pastures, grasslands, roadsides, railroad rights-of-way
Elevation: 300–2300(–2900) m

Distribution

V20-545-distribution-map.gif

Alta., B.C., Man., Ont., Sask., Colo., Idaho, Ill., Kans., Mich., Mont., Nebr., N.Dak., Oreg., S.Dak., Wash., Wyo.

Discussion

Variety villosa grows primarily in the Great Plains region; it is rare in Colorado, Idaho, Illinois, Kansas, and Oregon, and represented in Michigan and southern Ontario by chance introduction.

Selected References

None.

Lower Taxa

None.

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
paniculiform +, corymbiform +  and open +
indeterminate +  and determinate +
John C. Semple +
(Pursh) Shinners +
narrowly cuneate +  and attenuate +
Amellus villosus +
compound +  and simple +
2.8 cm28 mm <br />0.028 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1.25 cm12.5 mm <br />0.0125 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
oblanceolate;oblong rarely ovate or lanceolate-triangular +
3.5mm;7mm +
rugulose +  and muricate +
linear-oblanceolate +
reduced +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.85 cm8.5 mm <br />0.0085 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
not 2-lipped +  and actinomorphic +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
glabrate +  and glabrous +
ampliate +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
2.1 cm21 mm <br />0.021 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Alta. +, B.C. +, Man. +, Ont. +, Sask. +, Colo. +, Idaho +, Ill. +, Kans. +, Mich. +, Mont. +, Nebr. +, N.Dak. +, Oreg. +, S.Dak. +, Wash. +  and Wyo. +
300–2300(–2900) m +
strigose +  and stipitate-glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Silty, sandy loam or clay soils, chalk, or granitic soils, gravel soils, dry shale-limestone soils, red sandstone soils, travertine soils of sandhill prairies, pastures, grasslands, roadsides, railroad rights-of-way +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
narrowly cylindric;campanulate +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (2 cm20 mm <br />0.02 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
6.5mm;11mm +
0.85 cm8.5 mm <br />0.0085 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
sessile +  and petiolate +
cauline +  and basal +
0.075 cm0.75 mm <br />7.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
erect;spreading +
glabrate +  and glabrous +
lanceolate +, triangular +  and deltate +
reddish purple +
strigoso-ciliate +  and sparsely abundantly long-hispido-strigose +
undulate +
2-carpellate +
in 3 +  and 4 series +
4.7 cm47 mm <br />0.047 m <br /> (6.7 cm67 mm <br />0.067 m <br />) +
hispido-strigose +
1.2 cm12 mm <br />0.012 m <br /> (4.7 cm47 mm <br />0.047 m <br />) +
Flowering (May–)Jun–Oct(–Nov, south). +
triangular-lanceolate +
ovate-oblanceolate +
Field & Lab. +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
4 +  and 8 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
exalbuminous +
decumbent +  and erect +
1 +  and 50 +
16 cm160 mm <br />0.16 m <br /> (39 cm390 mm <br />0.39 m <br />) +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis mollis +
Heterotheca villosa var. villosa +
Heterotheca villosa +
variety +
40 cm400 mm <br />0.4 m <br /> (70 cm700 mm <br />0.7 m <br />) +
16 cm160 mm <br />0.16 m <br /> (40 cm400 mm <br />0.4 m <br />) +
tree +, shrub +  and subshrub +