Holocarpha macradenia

(de Candolle) Greene

Fl. Francisc. 4: 426. 1897.

EndemicConservation concern
Basionym: Hemizonia macradenia de Candolle in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 693. 1836
Treatment appears in FNA Volume 21. Treatment on page 288. Mentioned on page 287.

Plants 10–50 cm; stems notably stipitate-glandular. Heads borne singly or in glomerules or spiciform-glomerulate arrays. Involucres ± globose. Phyllaries each bearing ± 25 gland-tipped processes and minutely sessile or stipitate-glandular. Ray-florets 8–16. Disc-florets 40–90; anthers reddish to dark purple. 2n = 8.


Phenology: Flowering Jun–Nov.
Habitat: Grassy areas, clay soils
Elevation: 10–200 m

Discussion

Of conservation concern.

Holocarpha macradenia occurs on the coast from near Santa Cruz to central Monterey Bay area (most populations in the San Francisco Bay area are extirpated). Populations are reportedly highly interfertile; crosses to H. virgata have sometimes yielded moderately to highly fertile hybrids (the two species do not co-occur). J. Clausen (1951) and R. E. Palmer (1982) suggested that H. macradenia is most closely related to H. virgata; that hypothesis has been confirmed by molecular data (B. G. Baldwin, unpubl.).

Selected References

None.

Lower Taxa

None.
... more about "Holocarpha macradenia"
not tailed +, rounded +  and obtuse +
reddish +  and dark purple +
pale +  and dark +
connate +  and distinct +
subequal +
scarious +
usually ovate +  and lanceolate +
usually deltate +  and lanceolate +
umbelliform +, spiciform +, paniculiform +, glomerulate +  and corymbiform +
indeterminate +  and determinate +
Bruce G. Baldwin +  and John L. Strother +
(de Candolle) Greene +
decurrent +
Hemizonia macradenia +
compound +  and simple +
linear to linear-elliptic +
linear;oblanceolate +
winged;ribbed;winged;ribbed +
tuberculate +  and rugose +
barbellulate +  and barbellate +
hairy +  and glabrous +
fusiform +, clavate +, obpyramidal +, terete +, compressed +  and obcompressed +
fertile +  and bisexual +
fertile +  and bisexual +
10–200 m +
hirsute +  and strigose sericeous or villous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
spiciform-glomerulate +
Grassy areas, clay soils +
indeterminate +
homogamous +  and heterogamous +
each +  and sessile +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
petiolate +  and sessile +
alternate +  and opposite +
deltate +
entire +, serrate +  and serrulate +
2-carpellate +
persistent +
Flowering Jun–Nov. +
stipitate-glandular +
3 +  and 16 +
alternate +  and opposite +
Fl. Francisc. +
obcompressed +
flat;convex +
exalbuminous +
Endemic +  and Conservation concern +
stipitate-glandular +
appendaged +  and truncate +
hirsutulous +  and glabrous +
papillate +
Compositae +
Holocarpha macradenia +
Holocarpha +
species +
cylindric +
toothed +  and entire +
10 cm100 mm <br />0.1 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +