Castanea 9: 107. 1944.
Plants less than 3 cm. Stems with flagelliform branches absent. Stem and branch leaves 0.5–1.5 (–2) mm; basal laminal cells 20–27 × 9 µm; distal medial cells irregularly rhombic, 10 × 10 µm; apical cells roundedquadrate to short-oblong, 6–9 × 6 µm; basal marginal cells short-rhomboidal to fusiform-elongate, 9 × 8 µm, walls slightly pitted. Perichaetial inner leaves with base sheathing, margins entire to crenulate proximally, serrulate, apex obtuse, basal laminal cells rectangular, apical cells quadrate to rhombic.
Habitat: Shaded calcareous rock, cliffs or soil, hardwood-pine forests
Elevation: moderate elevations (200-500 m)
Ala., Mo., Tenn., Va., s Mexico, Asia (China), Asia (India), Asia (Japan)