Humulus lupulus var. lupuloides

E. Small

Syst. Bot. 3: 63. 1978 Humulus americanus Nuttall.

Endemic
Synonyms: Humulus americanus Nuttall
Treatment appears in FNA Volume 3.

Stems relatively pubescent at nodes, usually more than 15 hairs per 0.1 sq. mm at most pubescent portion (excluding angle of petiole with stem). Leaf-blades 10 cm or more usually having fewer than 5 lobes; smaller blades (ca. 5 cm) often with no more than 3 easily visible veins branching off midrib (excluding proximal branches); surfaces abaxially not conspicuously pubescent, usually fewer than 100 hairs per cm on length of medial midrib, more than 25 glands per 10 sq. mm between veins, hairs usually absent between veins.


Habitat: Moist thickets, woods, riverbanks
Elevation: 0-2000 m

Distribution

V3 963-distribution-map.gif

Alta., Man., N.B., Nfld. and Labr. (Nfld.), N.S., Ont., P.E.I., Que., Sask., Conn., Del., Ill., Ind., Iowa, Kans., Ky., Maine, Md., Mass., Mich., Minn., Mont., Nebr., N.H., N.J., N.Y., N.Dak., Ohio, Pa., R.I., S.Dak., Vt., Va., W.Va., Wis.

Discussion

Humulus lupulus var. lupuloides, defined as an intermediate, encompasses plants not falling into the other three varieties occurring in North America that are much more distinctive. It intergrades where it is sympatric with the introduced H. lupulus var. lupulus in southern Ontario, southern Quebec, and northeastern United States; with H. lupulus var. pubescens in midwestern United States; and with H. lupulus var. neomexicanus in the southern Canadian prairies and the Dakotas.

Selected References

None.

Lower Taxa

None.
compressed +  and ovoid +
Ernest Small +
E. Small +
10 cm100 mm <br />0.1 m <br /> (?) +
unlobed +, 3-7-lobed +  and cordate +
compound +  and staminate +
pendent +  and erect +
(10-)20-100+-flowered +
Alta. +, Man. +, N.B. +, Nfld. and Labr. (Nfld.) +, N.S. +, Ont. +, P.E.I. +, Que. +, Sask. +, Conn. +, Del. +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mont. +, Nebr. +, N.H. +, N.J. +, N.Y. +, N.Dak. +, Ohio +, Pa. +, R.I. +, S.Dak. +, Vt. +, Va. +, W.Va. +  and Wis. +
0-2000 m +
pistillate +, staminate +  and short-pedicellate +
paired +  and solitary +
whitish;yellowish +
loosely +  and covered +
transitional-bisexual +  and unisexual +
Moist thickets, woods, riverbanks +
2-branched +  and stalked +
straight +
pedunculate +  and staminate +
pendent +  and erect +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
3 cm30 mm <br />0.03 m <br /> (6 cm60 mm <br />0.06 m <br />) +
ovoid +  and oblong +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
unlobed +, lobed +  and cordate +
alternate +  and decussate +
compound +  and staminate +
pendent +  and erect +
mottled +  and brownish +
accrescent +
whitish +  and greenish +
hypogynous +
hypogynous +
filiform +
triangular +
resin-dotted and×or gland-dotted +
0 (?) +  and 100 (?) +
Humulus americanus +
Humulus lupulus var. lupuloides +
Humulus lupulus +
variety +
with soft pubescence +  and glabrous +
rhizomatous +  and taprooted +
perennial +  and annual +
rightward-twining +
600 cm6,000 mm <br />6 m <br /> (700 cm7,000 mm <br />7 m <br />) +