Humulus lupulus var. lupulus

IllustratedIntroduced
Treatment appears in FNA Volume 3.

Stems with sparse pubescence at nodes, usually fewer than 15 hairs per 0.1 sq. mm at most pubescent portion (excluding angle of petiole with stem). Leaf-blades usually with fewer than 20 hairs per cm on length of midrib, fewer than 25 glands per 10 sq. mm between veins.


Habitat: Roadsides, moist thickets, waste places, edges of woods
Elevation: 0-2000 m

Distribution

V3 1103-distribution-map.gif

Introduced; Man., N.B., N.S., Ont., P.E.I., Que., Calif., Conn., Del., Ill., Ind., Maine, Md., Mass., Mich., Mo., N.H., N.J., N.Y., Ohio, Pa., R.I., Vt., Va., W.Va., Wis.

Discussion

Humulus lupulus var. lupulus is indigenous to Europe. It has become established by introduction for ornament and for flavoring beer. Although not yet collected in all parts of the range, it should be expected, particularly as pistillate clones around abandoned homesteads. Humulus lupulus var. lupulusis reported from Oregon but no precise locality is known.

Selected References

None.

Lower Taxa

None.
compressed +  and ovoid +
Ernest Small +
Linnaeus +
3 cm30 mm <br />0.03 m <br /> (?) +  and 15 cm150 mm <br />0.15 m <br /> (?) +
unlobed +, 3-7-lobed +  and cordate +
compound +  and staminate +
pendent +  and erect +
(10-)20-100+-flowered +
Man. +, N.B. +, N.S. +, Ont. +, P.E.I. +, Que. +, Calif. +, Conn. +, Del. +, Ill. +, Ind. +, Maine +, Md. +, Mass. +, Mich. +, Mo. +, N.H. +, N.J. +, N.Y. +, Ohio +, Pa. +, R.I. +, Vt. +, Va. +, W.Va. +  and Wis. +
0-2000 m +
pistillate +, staminate +  and short-pedicellate +
paired +  and solitary +
whitish;yellowish +
loosely +  and covered +
transitional-bisexual +  and unisexual +
Roadsides, moist thickets, waste places, edges of woods +
2-branched +  and stalked +
straight +
pedunculate +  and staminate +
pendent +  and erect +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
3 cm30 mm <br />0.03 m <br /> (6 cm60 mm <br />0.06 m <br />) +
ovoid +  and oblong +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
unlobed +, lobed +  and cordate +
alternate +  and decussate +
compound +  and staminate +
pendent +  and erect +
mottled +  and brownish +
accrescent +
whitish +  and greenish +
hypogynous +
Illustrated +  and Introduced +
hypogynous +
filiform +
triangular +
resin-dotted and×or gland-dotted +
Humulus lupulus var. lupulus +
Humulus lupulus +
variety +
with soft pubescence +  and glabrous +
rhizomatous +  and taprooted +
perennial +  and annual +
rightward-twining +
600 cm6,000 mm <br />6 m <br /> (700 cm7,000 mm <br />7 m <br />) +