Huperzia porophila

(F. E. Lloyd & L. Underwood) Holub

Folia Geobot. Phytotax. 20: 76. 1985.

Basionym: Lycopodium porophilum F. E. Lloyd & L. Underwood Bull. Torrey Bot. Club 27(4): 150. 1900
Synonyms: Lycopodium lucidulum var. porophilum (F. E. Lloyd & L. Underwood) Clute Lycopodium selago var. porophilum (F. E. Lloyd & L. Underwood) Clute Urostachys lucidulus var. porophilus (F. Lloyd & L. Underwood) Nessel
Treatment appears in FNA Volume 2.

Shoots erect, determinate, occasionally indeterminate, 12–15 cm, clustered to short-decumbent, leaves of mature portion slightly smaller than leaves of juvenile portion; annual constrictions distinct to indistinct; juvenile growth erect. Leaves reflexed at base, ascending at stem apex (forming cluster) and spreading for most of stem length, sparse, yellow-green (juvenile portion) to yellow-green to green (mature portion), lustrous; largest leaves lanceolate with roughly parallel sides, 5–8 mm; smallest leaves triangular, widest at base, 3–6 mm; margins almost entire with low papillae or a few large teeth; stomates on both surfaces, few (1–25 per 1/2 leaf) on adaxial surface. Gemmiferous branchlets produced in 1–3 pseudowhorls at end of annual growth; gemmae 4–5 × 3–4 mm; lateral leaves 1–1.5 mm wide, acute, widest above middle. Spores 29–38 µm.


Habitat: On damp, shaded, acidic sandstone, rarely on shale or exposed sandstone
Elevation: 50–1200 m

Distribution

V2 537-distribution-map.gif

Ala., Ill., Ind., Iowa, Ky., Minn., Mo., Ohio, Pa., Tenn., Va., W.Va., Wis.

Discussion

Huperzia porophila occurs in several disjunct populations, most notably in the Driftless Area of Iowa, Minnesota, and Wisconsin. A sterile hybrid between H. porophila and H. lucidula, H. × bartleyi, occurs throughout the range of H. porophila. It reproduces vegetatively by gemmae and is common in habitats intermediate between those of the two parents. In Ohio the hybrid may occupy sites to which H. porophila would otherwise be restricted, thus reducing the sites available for colonization by H. porophila (A. W. Cusick 1987).

Selected References

None.

Lower Taxa

None.

"spreading" is not a number.

... more about "Huperzia porophila"
Warren H. Wagner Jr. +  and Joseph M. Beitel +
(F. E. Lloyd & L. Underwood) Holub +
Lycopodium porophilum +
Ala. +, Ill. +, Ind. +, Iowa +, Ky. +, Minn. +, Mo. +, Ohio +, Pa. +, Tenn. +, Va. +, W.Va. +  and Wis. +
50–1200 m +
linear;elliptic +
photosynthetic +
articulate +  and gemmiferous +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
according-to-seasonal +
On damp, shaded, acidic sandstone, rarely on shale or exposed sandstone +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
widest +, 1mm +  and 1.5mm +
pseudomonopodial +, dichotomous +, branching +, branched +  and simple +
flat +  and round +
monomorphic +
not imbricate +
yellow-green;yellow-green +
remote;dense +
mature +  and juvenile +
ascending;reflexed +
flattened +, lanceolate +  and oblanceolate +
slightly smaller +
aerial +  and subterranean +
juvenile +  and mature +
Folia Geobot. Phytotax. +
distinct +
emerging +
indeterminate +  and determinate +
0.2 cm2 mm <br />0.002 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
clustered +  and short-decumbent +
concave +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
plectostelic +  and actino +
axillary +  and adaxial +
protostelic +
subterranean +
stalked +  and sessile +
modified +  and unmodified +
photosynthetic +
small-grooved +
subterranean +
reticulate +, rugulate +  and pitted +
Lycopodium lucidulum var. porophilum +, Lycopodium selago var. porophilum +  and Urostachys lucidulus var. porophilus +
Huperzia porophila +
Huperzia +
species +
on rock +  and terrestrial +
erect +  and decumbent +