Isocoma acradenia var. bracteosa

(Greene) G. L. Nesom

Phytologia 70: 81. 1991.

Endemic
Basionym: Isocoma bracteosa Greene Leafl. Bot. Observ. Crit. 1: 170. 1906
Synonyms: Haplopappus acradenius subsp. bracteosus (Greene) H. M. Hall
Treatment appears in FNA Volume 20. Treatment on page 441. Mentioned on page 440.

Leaf-blades mostly 50–150 mm, margins of proximal usually shallowly toothed, of distal entire or serrulate. Involucres 7–8 mm. Phyllary apices spinulose-aristate. Florets 20–27.2n = 12.


Phenology: Flowering Aug–Oct.
Habitat: Salt flats
Elevation: 0–20 m

Discussion

In the limited zone of contact between vars. bracteosa and acradenia, intermediates are rare. The two differ in chromosome number and significantly in habitat (altitudes are essentially non-overlapping). Field studies may show them to warrant distinction at specific rank.

Selected References

None.

Lower Taxa

None.

"thick" is not a number.

not tailed +, rounded +  and obtuse +
distinct +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
spiciform +, racemiform +, paniculiform +  and corymbiform +
indeterminate +  and determinate +
Guy L. Nesom +
(Greene) G. L. Nesom +
decurrent +
Isocoma bracteosa +
5 cm50 mm <br />0.05 m <br /> (15 cm150 mm <br />0.15 m <br />) +
oblanceolate +  and narrowly obovate +
linear;oblanceolate or obovate +
rugulose +  and muricate +
not 2-lipped +  and actinomorphic +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
subterete +, terete +  and obpyramidal +
fertile +  and bisexual +
fertile +  and bisexual +
0–20 m +
gland-dotted;gland-dotted +
sericeous +, tomentose +, glabrous +, villous +  and hispidulous +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
Salt flats +
indeterminate +
heterogamous +  and homogamous +
stipitate-glandular +, hispidulous +  and glabrous +
each +  and sessile +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
obconic;turbinate or campanulate +
0.7 cm7 mm <br />0.007 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
deltate +
serrulate +  and entire +
lobed +, toothed +  and entire +
2-carpellate +
persistent +
Flowering Aug–Oct. +
oblong +  and elliptic-lanceolate +
spinulose-aristate +
styliferous +, neuter +  and pistillate +
bearing subulate enations +, hairy +  and bristly +
concave;usually flat;conic +
exalbuminous +
eglandular +  and few-branched +
resinous +
gland-dotted +
stipitate-glandular +, tomentose +, villous +, hispidulous +  and glabrous +
appendaged +  and truncate +
papillate +  and smooth +
Haplopappus acradenius subsp. bracteosus +
Isocoma acradenia var. bracteosa +
Isocoma acradenia +
variety +
goblet--shaped +
ampliate +
dark orange-resinous +
120 cm1,200 mm <br />1.2 m <br /> (150 cm1,500 mm <br />1.5 m <br />) +
20 cm200 mm <br />0.2 m <br /> (120 cm1,200 mm <br />1.2 m <br />) +
120 cm1,200 mm <br />1.2 m <br /> (150 cm1,500 mm <br />1.5 m <br />) +