Taxon	Illustrator ⠉
	Gratiola viscidula Leucospora multifida Limnophila sessiliflora	Barbara Alongi John Myers Barbara Alongi

Herbs, annual. Stems erect or ascending, glandular-villosulous. Leaves cauline, opposite or whorled; petiole present; blade not fleshy, not leathery, margins pinnatifid to bipinnatifid. Inflorescences axillary, flowers solitary; bracts absent. Pedicels present; bracteoles absent. Flowers bisexual; sepals 5, distinct, lanceolate to linear-lanceolate, calyx obscurely bilaterally symmetric, tubular; corolla pale blue to pale lavender, bilaterally symmetric, bilabiate, tubular, tube base not spurred or gibbous, throat not densely pilose internally, lobes 5, abaxial 3, adaxial 2; stamens 4, medially adnate to corolla, didynamous, filaments glabrous; staminode 0; ovary 2-locular, placentation axile; stigma cuneate. Fruits capsules, dehiscence septicidal. Seeds 50–100, white, parallel-ridged, ovoid, wings absent.

Distribution

c, e North America, n Mexico

Discussion

Species 2 (1 in the flora).

The second species of Leucospora, as circumscribed here, is L. coahuilensis Henrickson, which is known from Chihuahua, Coahuila, Durango, and Zacatecas, Mexico. It differs from L. multifida in its longer corollas, shorter pedicels, longer and more slender capsules, and smaller seeds (J. Henrickson 1989).

Based on a molecular phylogenetic study, D. Estes and R. L. Small (2008) suggested that Leucospora belongs to a clade that includes Bacopa, Gratiola, Mecardonia, Scoparia, and Stemodia, genera traditionally placed in Gratioleae. Relationships of some species, and the circumscriptions of some genera and the tribe itself, remain uncertain due to limited sampling. Leucospora was found to be sister to Stemodia verticillata (Miller) Hassler; Stemodia was paraphyletic, with the type species of that genus, S. maritima Linnaeus, included in a second, separate clade. B. L. Turner and C. C. Cowan (1993) included Leucospora in Stemodia based on morphological evidence.

Schistophragma, another genus possibly allied with Leucospora, has not been examined molecularly; it shares some morphological similarities with Leucospora. On the basis of morphology, J. Henrickson (1989) argued that Schistophragma should be submerged in Leucospora. Pending more complete taxon sampling for Gratioleae, Leucospora is retained here in its traditional sense.

Selected References

None.

Lower Taxa

Leucospora multifida