Leymus innovatus

(Beal) Pilg.
Common names: Downy ryegrass Boreal wildrye
Endemic
Synonyms: Elymus innovatus subsp. velutinus Elymus innovatus
Treatment appears in FNA Volume 24. Treatment on page 366.
Please click on the illustration for a higher resolution version.
Illustrator: Cindy Roché

Copyright: Utah State University

Plants sometimes cespitose, strongly rhizomatous. Culms 18-105 cm tall, 2-3 mm thick. Leaves exceeded by the spikes; sheaths glabrous or hairy, often most densely hairy in the collar region; auricles to 1.4 mm; ligules 0.1-0.5 mm; blades 2-6 mm wide, involute, abaxial surfaces scabridulous or smooth, adaxial surfaces scabrous, occasionally with scattered hairs to 1.5 mm, veins unequal, not crowded. Spikes 3-16 cm long, 8-20 mm thick, erect, usually well exserted, with 2-3 spikelets per node; internodes 4-6 mm, hairy throughout, edges with hairs to 2.5 mm. Spikelets 10-18 mm, with 3-7 florets. Glumes often unequal, sometimes absent, 2.5-12 mm long, 0.5-1 mm wide, hairy, stiff, keeled, the central portion thicker than the margins, bases not overlapping, tapering from near the base to the subulate apices, 0-1 (3) -veined, veins inconspicuous at midlength; calluses hairy; lemmas 7-12 mm, usually conspicuously villous or velutinous, occasionally glabrate, hairs 0.7-2.5 mm, awned, awns 2-4 mm; anthers 3.5-10 mm, dehiscent. 2n = 28, 56.

Distribution

Alta., B.C., Man., N.W.T., Ont., Sask., Yukon, Colo., Alaska, Mont., Wyo., S.Dak.

Discussion

Leymus innovatus is a North American species that grows in open woods and forests, riverbanks, open prairies, and rocky soils, and often in sandy, gravelly, or silty soils, primarily from northern Alaska to Hudson Bay, and south into the Black Hills region of Wyoming and South Dakota. Morphologically, the two subspecies show some overlap. Bowden recognized them in part because of their difference in ploidy level.

Selected References

None.

Key

1 Spikes 8-16 cm long, 8-15 mm wide; lemma hairs 0.7-2.5 mm long Leymus innovatus subsp. innovatus
1 Spikes 3-8 cm long, 15-20 mm wide; lemma hairs 1.5-2.5 mm long Leymus innovatus subsp. velutinus

"decumbent" is not a number."/2-33/4timesthelengthof" is not declared as a valid unit of measurement for this property.

... more about "Leymus innovatus"
smooth +  and scabridulous +
membranous +  and scarious +
0.35 cm3.5 mm <br />0.0035 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0-3(7)-veined +
straight +
midlength +
hairy +, glabrous +  and pilose +
subulate +
0 cm0 mm <br />0 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0 cm0 mm <br />0 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
Mary E. Barkworth +
(Beal) Pilg. +
0-3(7)-veined +
1 (?) +  and 3 (?) +
straight +
midlength +
glabrous +  and pilose +
tapering +, acuminate +, acute +  and keeled +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
keeled +  and rounded +
0-1(3)-veined +
not overlapping +
intravaginal +, extravaginal +, branching +  and basal +
tapering +
swelling +
not evident +
cross +, linear +  and narrowly lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
spikelike +
Downy ryegrass +  and Boreal wildrye +
not branching +
18 cm180 mm <br />0.18 m <br /> (105 cm1,050 mm <br />1.05 m <br />) +
10 cm100 mm <br />0.1 m <br /> (350 cm3,500 mm <br />3.5 m <br />) +
not woody +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Alta. +, B.C. +, Man. +, N.W.T. +, Ont. +, Sask. +, Yukon +, Colo. +, Alaska +, Mont. +, Wyo. +  and S.Dak. +
liquid +, soft +  and hard +
capillary +
pistillate +  and staminate +
3 +  and 7 +
laterally compressed +  and terete +
shorter than to longer than the adjacent florets +
2.5mm +  and 12mm +
subtending +
unequal +
0.5mm +  and 1mm +
uncinate +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
solid +  and hollow +
15 cm150 mm <br />0.15 m <br /> (?) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0-3(7)-veined +
glabrous +  and pilose +
tapering +, acuminate +, acute +  and keeled +
distributed +
glabrate +, velutinous +  and villous +
keeled +  and rounded +
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
stiffly membranous +  and coriaceous +
truncate +  and rounded +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
membranous +
inconspicuous +
fleshy +  and membranous +
concealed +  and prominent +
2 +  and 1 +
hairy +  and glabrous +
well-developed +
slightly shorter than to slightly longer than the lemmas +
slightly longer +
subequal +
appressed +  and ascending +
0 cm0 mm <br />0 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
dry +  and fleshy +
spikelike +
hairy +  and glabrous +
3 cm30 mm <br />0.03 m <br /> (?) +  and 16 cm160 mm <br />0.16 m <br /> (?) +
0.8 cm8 mm <br />0.008 m <br /> (?) +  and 2 cm20 mm <br />0.02 m <br /> (?) +
pedicellate +  and sessile +
bisexual +  and sterile +
terete +  and compressed +
1 cm10 mm <br />0.01 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1 +  and 3 +
simple +  and compound +
2 (?) +  and 3 (?) +
not papillate +
Elymus innovatus subsp. velutinus +  and Elymus innovatus +
Leymus innovatus +
species +
awnlike +
membranous +
0-3(7)-veined +
awned +  and ribbed +
not crowded +
midlength +
inconspicuous +
glabrous +  and pilose +
tapering +, acuminate +, acute +  and keeled +
unequal +
0 cm0 mm <br />0 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
plant +  and cespitose +
aquatic +  and terrestrial +