Minuartia nuttallii

(Pax) Briquet

Annuaire Conserv. Jard. Bot. Genève 13–14: 385. 1911.

Common names: Nuttall’s sandwort
Endemic
Basionym: Arenaria nuttallii Pax Bot. Jahrb. Syst. 18: 30. 1893
Synonyms: Minuopsis nuttallii (Pax) W. A. Weber
Treatment appears in FNA Volume 5. Treatment on page 129. Mentioned on page 119, 124, 133, 134.

Plants perennial, mat-forming. Taproots thickened, woody; crown, many-branched, woody; rhizomes and trailing stems to 60 cm. Stems ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering-stems 0.2–2 times as long as leaves. Leaves tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm; blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular; axillary leaves present proximally to throughout. Inflorescences (3–) 6–30-flowered, open cymes; bracts lanceolate to subulate, usually scarious. Pedicels 0.2–2 cm, stipitate-glandular. Flowers: hypanthium disc-shaped; sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6 (–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular; petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire. Capsules on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals. Seeds reddish-brown to dark-brown, oblongelliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate; tubercles low-rounded.

Distribution

V5 264-distribution-map.gif

Alta., B.C., Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Varieties 4 (4 in the flora).

Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit.

Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis).

Selected References

None.

Key

1 Leaf blade apices spinecent; sepal apices spinescent > 2
1 Leaf blade apices acute to acuminate, somewhat navicular; sepal apices acuminate > 3
2 Leaves prominently arcuate-spreading, blade 10-20 mm; sepals (1-)3-veined Minuartia nuttallii var. fragilis
2 Leaves appressed to occasionally arcuate- spreading, blade 5-7 mm; sepals 1(-3)-veined Minuartia nuttallii var. gracilis
3 Sepals lanceolate to narrowly so; petals 0.5-1.2 times as long as sepals Minuartia nuttallii var. nuttallii
3 Sepals ovate to lanceolate; petals 1.1-1.6 times as long as sepals Minuartia nuttallii var. gregaria
... more about "Minuartia nuttallii"
not hooded +  and spinescent +
green +  and purple +
stipitate-glandular +
rounded;acute;acuminate +
Richard K. Rabeler +, Ronald L. Hartman +  and Frederick H. Utech +
(Pax) Briquet +
Arenaria nuttallii +
straight +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
broadly lanceolate +  and linear +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
lanceolate +  and subulate +
reduced +
5 cm50 mm <br />0.05 m <br /> (?) +
Nuttall’s sandwort +
many-branched +
axillary +  and terminal +
dish--shaped +
expanded +
Alta. +, B.C. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
straight +  and curved +
central +  and peripheral +
axillary +  and terminal +
inconspicuous +
unisexual +  and bisexual +
stalked +  and simple +
disc--shaped +  and dish +
disc--shaped +
(3-)6-30-flowered +
0.2-2 times as long as leaves +
sessile +  and petiolate +
connate +
tightly appressed;spreading +
swollen +
3 +  and 5 +
crassinucellate +  and bitegmic +
campylotropous +
reflexed;erect;arcuate-spreading +
stipitate-glandular +
0.2 cm2 mm <br />0.002 m <br /> (2 cm20 mm <br />0.02 m <br />) +
lilac +, pink +  and white +
fugacious +
0.5-1.6 times as long as sepals +
obovate +
Annuaire Conserv. Jard. Bot. Genève +
0 cm0 mm <br />0 m <br /> (60 cm600 mm <br />0.6 m <br />) +
reddish-brown;dark-brown +
tuberculate +
oblongelliptic +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
persistent +
distinct +
narrowly lanceolate;lanceolate or ovate +
not enlarging +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
branched +  and simple +
ascending;erect +
glandular-hairy +
2 cm20 mm <br />0.02 m <br /> (20 cm200 mm <br />0.2 m <br />) +
[30 +  and 3 +
papillate +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (?) +  and 0.02 cm0.2 mm <br />2.0e-4 m <br /> (?) +
distinct +
filiform +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Minuopsis nuttallii +
Minuartia nuttallii +
Minuartia +
species +
divided +
low-rounded +
10 +  and 8 +
erect +  and recurved +
rhizomatous +  and taprooted +
plant +  and mat-forming +
pubescence of simple hairs or stalked glands +  and glabrous +
23 +, 15 +, 14 +, 13 +, 12 +, 10 +, 9 +  and 7 +