genusNajas

Najas marina

Linnaeus

Sp. Pl. 2: 1015. 1753..

Common names: Spiny naiad
Illustrated
Synonyms: Najas gracilis (Morong) Small
Treatment appears in FNA Volume 22.

Stems branched distally, 6–45 cm × 0.5–4 mm; internodes 0.3–11 cm, usually with prickles. Leaves spreading to ascending with age, 0.5–3.9 cm, stiff in age; sheaths 2–4.4 mm wide, apex acute; blade 0.4–4.5 mm wide, margins coarsely serrate, teeth 8–13 per side, apex acute, with 1 tooth, teeth multicellular; midvein with prickles abaxially. Flowers 1 per axil, staminate and pistillate on different plants. Staminate flowers in distal to proximal axils, 1.7–3 mm; involucral beaks 2-lobed, 0.3–0.7 mm; anthers 4-loculed, 1.7–3 mm. Pistillate flowers in distal to proximal axils, 2.5–5.7 mm; styles 1.2–1.7 mm; stigmas 3-lobed. Seeds not recurved, reddish-brown, ovoid, 2.2–4.5 × 1.2–2.2 mm, apex with style situated at center; testa dull, 10–15 cell-layers thick, pitted; areoles irregularly arranged, not in distinctive rows, not ladderlike, 3–4-angled, longer than broad, end walls slightly raised. 2n = 12 (Europe).


Phenology: Flowering summer–fall.
Habitat: Brackish or highly alkaline waters of ponds and lakes
Elevation: 0–1000 m

Distribution

V22 492-distribution-map.jpg

Ariz., Calif., Fla., Ind., Mich., Minn., Nev., N.Y., N.Dak., Ohio, Pa., S.Dak., Tex., Utah, Wis., Mexico, West Indies, Central America (El Salvador), Central America (Panama), South America, Eurasia

Discussion

With its prickly internodes and prickles along the abaxial surface of the leaves, Najas marina is the easiest of our Najas to recognize. Over its entire range, the species displays considerable morphologic variability (L. Triest et al. 1986), giving cause for ten subspecies to be recognized (L. Triest 1988). In North America, however, variability is relatively minor, so I am recognizing the taxon at the specific level only (R. R. Haynes 1979). The species should be studied over its entire range, utilizing a variety of approaches, to determine adequately what, if any, infraspecific taxa should be recognized.

Lower Taxa

None.

"elongating" is not a number.

... more about "Najas marina"
0.17 cm1.7 mm <br />0.0017 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
longer than broad +
3-4-angled +  and not ladderlike +
Robert R. Haynes +
Linnaeus +
involucral +
2-lobed +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
armed +  and 1-veined +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
in 15-60 longitudinal rows +  and arranged +
sharp-tipped +
Spiny naiad +
short-pedunculate +  and sessile +
Ariz. +, Calif. +, Fla. +, Ind. +, Mich. +, Minn. +, Nev. +, N.Y. +, N.Dak. +, Ohio +, Pa. +, S.Dak. +, Tex. +, Utah +, Wis. +, Mexico +, West Indies +, Central America (El Salvador) +, Central America (Panama) +, South America +  and Eurasia +
0–1000 m +
pistillate +  and staminate +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.57 cm5.7 mm <br />0.0057 m <br />) +
Brackish or highly alkaline waters of ponds and lakes +
0.3 cm3 mm <br />0.003 m <br /> (11 cm110 mm <br />0.11 m <br />) +
light green +  and brown +
red-purple +  and purple +
whorled +, appearing +  and opposite +
submersed +
spreading;ascending +
0.5 cm5 mm <br />0.005 m <br /> (3.9 cm39 mm <br />0.039 m <br />) +
serrate to minutely +
Flowering summer–fall. +
not stipitate +
stuckey1985a +, triest1986a +  and viinikka1973a +
distinctive +
15 +  and 60 +
auriculate +  and not auriculate +
reddish-brown +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
not recurved +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
not persisting +
variously--shaped +
2mm;4.4mm +
Illustrated +
6 cm60 mm <br />0.06 m <br /> (45 cm450 mm <br />0.45 m <br />) +
slender +
0.5mm;4mm +
2-4-branched +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
Najas gracilis +
Najas marina +
species +
10 +  and 15 +
multicellular +
caulescent +  and not rhizomatous +
perennial +  and annual +