Oenothera cavernae

Munz

Leafl. W. Bot. 3: 50. 1941.

Treatment appears in FNA Volume 10.

Herbs winter or spring annual, acaulescent or short-caulescent, glandular puberulent, some­times also sparsely hirsute; from a taproot. Stems (when present) 1–several, ascending, usually unbranched, 2–4 cm. Leaves primarily in a basal rosette, sometimes also cauline, (0.5–) 2.5–13 (–19.5) × (0.2–) 0.6–2.3 (–2.7) cm; petiole 0.5–5.2 cm; blade oblanceolate to elliptic-oblanceo­late (in some exceptionally large leaves), margins lyrate-pinnatifid to subentire (in very small ones), apex usually rounded, rarely acute. Flowers 1–3 (–10) per stem opening per day near sunset, without noticeable scent; buds sometimes ± recurved before anthesis; floral-tube (20–) 30–37 (–47) mm; sepals 4.5–12 mm; petals white, fading pale-pink, (6.5–) 8–20 (–25) mm; filaments 5.2–7.5 (–12) mm, anthers (1.4–) 3–4.5 (–6) mm; style (24–) 35–45 (–56) mm, stigma surrounded by anthers at anthesis. Capsules falcate (especially before maturity), ellipsoid-ovoid to ovoid, obtusely 4-angled, 12–38 × 6–14 mm, tapering to a sterile beak 2–8 mm, dehis­cent to 1/2 their length, valve margins with a sinuate ridge or 8–20 nearly distinct tubercles; pedicel 0–10 mm. Seeds usually numerous, sometimes as few as 5, in 2 adjacent rows per locule, obovoid, 2.5–3.1 × 1.1–1.4 mm, embryo 1/2 of seed volume, surface minutely papillose to reticu­late; seed collar without membrane, producing a large empty cavity, margin irregularly sinuate. 2n = 14.


Phenology: Flowering Mar–May.
Habitat: Exposed calcareous slopes, crevices in limestone, dolomite, or loose talus, sandy arroyos, sandstone, granitic crevices, volcanic cinders in Mojave Desert or Great Basin scrub communities, rarely in arid juniper woodlands.
Elevation: 400–1700 m.

Discussion

Oenothera cavernae is known from the Arrow Canyon, Las Vegas, and Sheep ranges and the low hills near Arden and Sloan in Clark County, Nevada, eastward along the Grand Canyon to the vicinity of Page, Arizona, and perhaps Washington County, Utah and formerly in Glenn Canyon, and more recently collected in eastern San Bernardino County, Cali­fornia (eastern Clark Mountain Range, and the base of range in Ivanpah Valley). W. L. Wagner et al. (1985) deter­mined O. cavernae to be self-compatible and autogamous.

Selected References

None.

Lower Taxa

None.
... more about "Oenothera cavernae"
0.45 cm4.5 mm <br />0.0045 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
acute +  and rounded +
Warren L. Wagner +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
pedicel-like +
Pachylophus +
0.2 cm2 mm <br />0.002 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
indehiscent +  and loculicidal +
subentire +, serrate +, usually coarsely dentate +  and pinnatifid +
oblanceolate +
quadrangular +
indehiscent +  and dehiscent +
1.2 cm12 mm <br />0.012 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
tapering +, 4-angled +, ellipsoid-ovoid +  and ovoid +
0.6 cm6 mm <br />0.006 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Ariz. +, Calif. +  and Nev. +
400–1700 m. +
whitish +  and green +
exfoliating +
0.75 cm7.5 mm <br />0.0075 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
basifixed +  and versatile +
0.52 cm5.2 mm <br />0.0052 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
3.7 cm37 mm <br />0.037 m <br /> (4.7 cm47 mm <br />0.047 m <br />) +
hispid +, lanate +  and glabrous +
3 cm30 mm <br />0.03 m <br /> (3.7 cm37 mm <br />0.037 m <br />) +
zygomorphic +  and actinomorphic +
unscented +
1 +  and 3 +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Exposed calcareous slopes, crevices in limestone, dolomite, or loose talus, sandy arroyos, sandstone, granitic crevices, volcanic cinders in Mojave Desert or Great Basin scrub communities, rarely in arid juniper woodlands. +
short-caulescent +  and acaulescent +
suffrutescent +
hirsute +  and puberulent +
spring +  and winter +
nodding +  and erect +
13 cm130 mm <br />0.13 m <br /> (19.5 cm195 mm <br />0.195 m <br />) +
26 cm260 mm <br />0.26 m <br /> (36 cm360 mm <br />0.36 m <br />) +
2.3 cm23 mm <br />0.023 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
deciduous +
2.5 cm25 mm <br />0.025 m <br /> (13 cm130 mm <br />0.13 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (26 cm260 mm <br />0.26 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
sinuate +, lyrate-pinnatifid +  and subentire +
pedicellate +  and sessile +
1 +  and 8 +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
fading pale-pink +  and white +
obcordate +  and obovate +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (5.2 cm52 mm <br />0.052 m <br />) +
Flowering Mar–May. +
Leafl. W. Bot. +
distinct +
sinuate +
basal +  and cauline +
clavate +  and globose +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.31 cm3.1 mm <br />0.0031 m <br />) +
5 +  and numerous +
sculptured +  and smooth +
obovoid +
reduced +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
in pairs +  and separating +
purple +, pink +, red +  and green +
deciduous +
0.45 cm4.5 mm <br />0.0045 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
1 +  and several +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
petiolate +  and sessile +
intrapetiolar +
4.5 cm45 mm <br />0.045 m <br /> (5.6 cm56 mm <br />0.056 m <br />) +
pubescent +  and glabrous +
3.5 cm35 mm <br />0.035 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
papillose +
Oenothera +  and Oenothera subg. Pachylophus +
Oenothera cavernae +
Oenothera sect. Pachylophus +
species +
3(-5)-aperturate +
pedicellate +  and sessile +
woody +  and hard +
smooth +  and erose +
papillate +