Oenothera cespitosa subsp. macroglottis

(Rydberg) W. L. Wagner, Stockhouse & W. M. Klein

Ann. Missouri Bot. Gard. 70: 195. 1983. (as caespitosa)

Basionym: Pachylophus macroglottis Rydberg Bull. Torrey Bot. Club 30: 259. 1903
Synonyms: Oenothera cespitosa var. macroglottis (Rydberg) Cronquist P. hirsutus Rydberg
Treatment appears in FNA Volume 10.

Herbs acaulescent or short-caulescent, hirsute and glan­dular puberulent, or glabrous. Stems (if present), usually unbranched, rarely with 1–several short laterals, 4–8 cm. Leaves (6.8–) 9.5–23 (–32) × (1.3–) 2.4–4.5 (–6.5) cm; petiole (3–) 4–11 (–14) cm; blade oblanceolate to spatulate, margins often undulate, usually regularly to irreg­ularly dentate, rarely coarsely and irregularly pinnately lobed. Flowers: floral-tube (45–) 75–110 (–153) mm; sepals (22–) 30–45 (–50) mm; petals fading pink to pale rose, (21–) 35–43 (–50) mm; filaments (16–) 19–28 (–35) mm, anthers (10–) 12–17 mm; style 85–180 mm. Capsules somewhat curved, lanceoloid-cylindrical to cylindrical, symmetrical throughout, sometimes slightly flattened on one side at base, (17–) 25–45 (–56) × 5–8 mm, valve margins with conspicuous, nearly smooth to irregular undulate ridge; pedicel 2–7 mm. Seeds narrowly obovoid, 2.5–3 × 1–1.4 mm, embryo 1/2 of seed volume, surface minutely papillose to reticulate; seed collar forming narrow slit above raphe with a slightly sunken membrane, margin entire or obscurely sinuate distally. 2n = 14, 28.


Phenology: Flowering May–Jul(–Sep).
Habitat: Open, igneous rocky slopes, talus, roadcuts, open or shaded and sandy or gravelly sites along streams, rarely on shale, in upper pinyon-juniper woodlands, Gambel oak scrub, ponderosa pine forests, ponderosa pine-Douglas fir forests, spruce-fir-lodgepole pine forests.
Elevation: 2000–3100 m.

Discussion

Selected References

None.

Lower Taxa

None.
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
acuminate +, usually acute +  and rounded +
Warren L. Wagner +
(Rydberg) W. L. Wagner, Stockhouse & W. M. Klein +
4.5 cm45 mm <br />0.045 m <br /> (5.6 cm56 mm <br />0.056 m <br />) +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
2.5 cm25 mm <br />0.025 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
pedicel-like +
5mm;8mm +
Pachylophus macroglottis +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
indehiscent +  and loculicidal +
subentire +, serrate +, usually coarsely dentate +  and pinnatifid +
oblanceolate +  and spatulate +
recurved +  and erect +
quadrangular +
5 cm50 mm <br />0.05 m <br /> (6.8 cm68 mm <br />0.068 m <br />) +
indehiscent +  and dehiscent +
1.3 cm13 mm <br />0.013 m <br /> (5 cm50 mm <br />0.05 m <br />) +
flattened +, lanceoloid-cylindrical +  and cylindrical +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
depressed +  and flat +
Colo. +, N.Mex. +, Utah +  and Wyo. +
2000–3100 m. +
whitish +  and green +
exfoliating +
2.8 cm28 mm <br />0.028 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
basifixed +  and versatile +
1.9 cm19 mm <br />0.019 m <br /> (2.8 cm28 mm <br />0.028 m <br />) +
11 cm110 mm <br />0.11 m <br /> (15.3 cm153 mm <br />0.153 m <br />) +
hispid +, lanate +  and glabrous +
7.5 cm75 mm <br />0.075 m <br /> (11 cm110 mm <br />0.11 m <br />) +
zygomorphic +  and actinomorphic +
unscented +
1 +  and 4 +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Open, igneous rocky slopes, talus, roadcutOpen, igneous rocky slopes, talus, roadcuts, open or shaded and sandy or gravelly sites along streams, rarely on shale, in upper pinyon-juniper woodlands, Gambel oak scrub, ponderosa pine forests, ponderosa pine-Douglas fir forests, spruce-fir-lodgepole pine forests.orests, spruce-fir-lodgepole pine forests. +
short-caulescent +  and acaulescent +
perennial +
suffrutescent +
glabrous +, puberulent +  and hirsute +
nodding +  and erect +
23 cm230 mm <br />0.23 m <br /> (32 cm320 mm <br />0.32 m <br />) +
26 cm260 mm <br />0.26 m <br /> (36 cm360 mm <br />0.36 m <br />) +
4.5 cm45 mm <br />0.045 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
deciduous +
9.5 cm95 mm <br />0.095 m <br /> (23 cm230 mm <br />0.23 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (26 cm260 mm <br />0.26 m <br />) +
2.4 cm24 mm <br />0.024 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
sinuate +, entire +, lobed +  and undulate +
pedicellate +  and sessile +
1 +  and 8 +
4 cm40 mm <br />0.04 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
4.3 cm43 mm <br />0.043 m <br /> (5 cm50 mm <br />0.05 m <br />) +
fading pink +  and pale rose +
obcordate +  and obovate +
3.5 cm35 mm <br />0.035 m <br /> (4.3 cm43 mm <br />0.043 m <br />) +
11 cm110 mm <br />0.11 m <br /> (14 cm140 mm <br />0.14 m <br />) +
4 cm40 mm <br />0.04 m <br /> (11 cm110 mm <br />0.11 m <br />) +
Flowering May–Jul(–Sep). +
Ann. Missouri Bot. Gard. +
smooth to irregular +
conspicuous +
undulate +
basal +  and cauline +
sinuate +
clavate +  and globose +
depressed +  and flat +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
sculptured +  and smooth +
obovoid +
reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
in pairs +  and separating +
4.5 cm45 mm <br />0.045 m <br /> (5 cm50 mm <br />0.05 m <br />) +
purple +, pink +, red +  and green +
deciduous +
3 cm30 mm <br />0.03 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
decumbent +  and ascending +
4 cm40 mm <br />0.04 m <br /> (8 cm80 mm <br />0.08 m <br />) +
petiolate +  and sessile +
intrapetiolar +
18 cm180 mm <br />0.18 m <br /> (18.5 cm185 mm <br />0.185 m <br />) +
pubescent +  and glabrous +
8.5 cm85 mm <br />0.085 m <br /> (18 cm180 mm <br />0.18 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
minutely papillose +  and reticulate +
Oenothera cespitosa var. macroglottis +  and P. hirsutus +
Oenothera cespitosa subsp. macroglottis +
Oenothera cespitosa +
subspecies +
3(-5)-aperturate +
pedicellate +  and sessile +
woody +  and hard +
smooth +  and erose +
papillate +