Oenothera cespitosa subsp. navajoensis

W. L. Wagner, Stockhouse & W. M. Klein

Monogr. Syst. Bot. Missouri Bot. Gard. 12: 66, fig. 104. 1985. (as caespitosa)

Synonyms: Oenothera cespitosa var. navajoensis (W. L. Wagner, Stockhouse & W. M. Klein) Cronquist Pachylophus caulescens Rydberg
Treatment appears in FNA Volume 10.
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Herbs caulescent or acaulescent, moderately to densely crinkly-villous (often appearing shaggy), and glandular puberulent. Stems (if present) unbranched to few-branched, (0–) 10–25 cm. Leaves (3.5–) 4–13 (–16) × (0.7–) 1–3.2 cm; petiole (1.3–) 1.7–10 (–12) cm; blade oblanceolate to rhombic-obovate, margins often coarsely and irregularly dentate or serrate, sometimes pinnately lobed, often with several larger lobes near blade base. Flowers: buds often recurved when young; floral-tube (35–) 40–70 (–80) mm; sepals 22–27 (–32) mm; petals fading pink to light rose, (25–) 28–32 (–34) mm; filaments 11–15 (–17) mm, anthers 9–12 mm; style (50–) 59–85 (–96) mm. Capsules straight, oblong-lanceoloid, base asymmetrical, 13–35 (–40) × 5–6 mm; valve margins with a low sinuate ridge to 8–15 small, nearly distinct tubercles; pedicel 1–3 mm. Seeds narrowly obovoid, 2.1–2.6 × 1.1–1.3 mm, embryo 1/2 of seed volume, surface minutely papillose; seed collar forming a narrow slit above raphe with a slightly sunken membrane, mar­gin entire or obscurely sinuate distally. 2n = 14, 28.


Phenology: Flowering (Apr–)May–Jun.
Habitat: Colorado Plateau region, forming small colonies on loose or compacted soil derived from clay, shale, fine-textured sandstone, or gypsum, on slopes and along small drainage patterns, often around harvester ant mounds, arroyos in somewhat sandy or gravelly soil, in shrubby commu­nities dominated by Atriplex confertifolia, A. corrugata, A. cuneata, Artemisia spinescens, Coleogyne ramosissima, Frankenia jamesii, Hilaria jamesii, with big sagebrush scrub or sage-grasslands, rarely in lower pinyon-juniper woodlands.
Elevation: 1100–1900(–2100) m.

Distribution

Ariz., Colo., N.Mex., Utah.

Discussion

Selected References

None.

Lower Taxa

None.
1.7 cm17 mm <br />0.017 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
acuminate +, usually acute +  and rounded +
Warren L. Wagner +
W. L. Wagner, Stockhouse & W. M. Klein +
3.5 cm35 mm <br />0.035 m <br /> (4 cm40 mm <br />0.04 m <br />) +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
1.3 cm13 mm <br />0.013 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
pedicel-like +
5mm;6mm +
Pachylophus +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
indehiscent +  and loculicidal +
subentire +, serrate +, usually coarsely dentate +  and pinnatifid +
oblanceolate +  and rhombic-obovate +
quadrangular +
5 cm50 mm <br />0.05 m <br /> (6.8 cm68 mm <br />0.068 m <br />) +
indehiscent +  and dehiscent +
1.3 cm13 mm <br />0.013 m <br /> (5 cm50 mm <br />0.05 m <br />) +
oblong-lanceoloid +
0.4 cm4 mm <br />0.004 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
depressed +  and flat +
Ariz. +, Colo. +, N.Mex. +  and Utah. +
1100–1900(–2100) m. +
whitish +  and green +
exfoliating +
1.5 cm15 mm <br />0.015 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
basifixed +  and versatile +
1.1 cm11 mm <br />0.011 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
7 cm70 mm <br />0.07 m <br /> (8 cm80 mm <br />0.08 m <br />) +
hispid +, lanate +  and glabrous +
4 cm40 mm <br />0.04 m <br /> (7 cm70 mm <br />0.07 m <br />) +
zygomorphic +  and actinomorphic +
unscented +
1 +  and 4 +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Colorado Plateau region, forming small colColorado Plateau region, forming small colonies on loose or compacted soil derived from clay, shale, fine-textured sandstone, or gypsum, on slopes and along small drainage patterns, often around harvester ant mounds, arroyos in somewhat sandy or gravelly soil, in shrubby communities dominated by Atriplex confertifolia, A. corrugata, A. cuneata, Artemisia spinescens, Coleogyne ramosissima, Frankenia jamesii, Hilaria jamesii, with big sagebrush scrub or sage-grasslands, rarely in lower pinyon-juniper woodlands. rarely in lower pinyon-juniper woodlands. +
acaulescent +  and caulescent +
perennial +
suffrutescent +
puberulent +  and crinkly-villous +
nodding +  and erect +
13 cm130 mm <br />0.13 m <br /> (16 cm160 mm <br />0.16 m <br />) +
26 cm260 mm <br />0.26 m <br /> (36 cm360 mm <br />0.36 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
deciduous +
4 cm40 mm <br />0.04 m <br /> (13 cm130 mm <br />0.13 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (26 cm260 mm <br />0.26 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
serrate +  and dentate +
pedicellate +  and sessile +
1 +  and 8 +
4 cm40 mm <br />0.04 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
3.2 cm32 mm <br />0.032 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
fading pink +  and light rose +
obcordate +  and obovate +
2.8 cm28 mm <br />0.028 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
10 cm100 mm <br />0.1 m <br /> (12 cm120 mm <br />0.12 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (10 cm100 mm <br />0.1 m <br />) +
Flowering (Apr–)May–Jun. +
Monogr. Syst. Bot. Missouri Bot. Gard. +
sinuate +
basal +  and cauline +
sinuate +
clavate +  and globose +
sinuate +  and entire +
0.21 cm2.1 mm <br />0.0021 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
sculptured +  and smooth +
obovoid +
reduced +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
in pairs +  and separating +
2.7 cm27 mm <br />0.027 m <br /> (3.2 cm32 mm <br />0.032 m <br />) +
purple +, pink +, red +  and green +
deciduous +
2.2 cm22 mm <br />0.022 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
unbranched +  and few-branched +
0 cm0 mm <br />0 m <br /> (10 cm100 mm <br />0.1 m <br />) +
decumbent +  and ascending +
10 cm100 mm <br />0.1 m <br /> (25 cm250 mm <br />0.25 m <br />) +
petiolate +  and sessile +
intrapetiolar +
8.5 cm85 mm <br />0.085 m <br /> (9.6 cm96 mm <br />0.096 m <br />) +
pubescent +  and glabrous +
5.9 cm59 mm <br />0.059 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
papillose +
Oenothera cespitosa var. navajoensis +  and Pachylophus caulescens +
Oenothera cespitosa subsp. navajoensis +
Oenothera cespitosa +
subspecies +
3(-5)-aperturate +
pedicellate +  and sessile +
woody +  and hard +
smooth +  and erose +
papillate +