Oenothera deltoides subsp. piperi

(Munz) W. M. Klein

Aliso 5: 180. 1962.

Basionym: Oenothera deltoides var. piperi Munz Amer. J. Bot. 18: 314. 1931
Synonyms: O. trichocalyx var. piperi (Munz) Jepson
Treatment appears in FNA Volume 10.

Herbs annual, usually villous, hairs relatively long, curly, especially distally and on buds, sometimes glabrous; from a taproot. Stems: central stem thickened proximally, unbranched or with several lateral, ascending to decumbent branches, 3–30 (–40) cm. Leaves basal and cauline; blade rhombic, becoming lanceolate distally, margins deeply sinuate-dentate to pinnatifid. Flowers: buds fluted or strongly quadran­gular in distal 1/2, without free tips; sepals 13–22 (–27) mm; petals 15–25 (–30) mm. Capsules 15–25 (–30) × 3–5 mm. 2n = 14.


Phenology: Flowering (Mar–)Jun–Jul(–Sep).
Habitat: Sandy soil or dunes in Great Basin Desert with Artemisia, Ericameria, or Sarcobatus.
Elevation: 900–1900 m.

Discussion

W. M. Klein (1964) determined subsp. piperi to be self-incompatible. It occurs in the northern part of the range of O. deltoides, from northeastern California to southern Oregon and the western half of Nevada.

Selected References

None.

Lower Taxa

None.
0.5 cm5 mm <br />0.005 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
1.5mm +  and 5mm +
Warren L. Wagner +
(Munz) W. M. Klein +
5 cm50 mm <br />0.05 m <br /> (25 cm250 mm <br />0.25 m <br />) +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
pedicel-like +
Oenothera deltoides var. piperi +
indehiscent +  and loculicidal +
sinuate-dentate +  and pinnatifid denticulate subentire or entire +
lanceolate +  and rhombic +
40 cm400 mm <br />0.4 m <br /> (100 cm1,000 mm <br />1 m <br />) +
ascending +  and decumbent +
slender +
10 cm100 mm <br />0.1 m <br /> (40 cm400 mm <br />0.4 m <br />) +
strongly +  and fluted +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
straight +  and curved +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
tapering +, cylindrical +  and slightly 4-angled +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
12 cm120 mm <br />0.12 m <br /> (18 cm180 mm <br />0.18 m <br />) +
4 cm40 mm <br />0.04 m <br /> (12 cm120 mm <br />0.12 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (4 cm40 mm <br />0.04 m <br />) +
with several lateral , ascending to decumbent branches +  and unbranched +
30 cm300 mm <br />0.3 m <br /> (40 cm400 mm <br />0.4 m <br />) +
3 cm30 mm <br />0.03 m <br /> (30 cm300 mm <br />0.3 m <br />) +
Calif. +, Nev. +  and Oreg. +
900–1900 m. +
pink +  and white +
exfoliating +
basifixed +  and versatile +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
4 cm40 mm <br />0.04 m <br /> (5 cm50 mm <br />0.05 m <br />) +
hispid +, lanate +  and glabrous +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
zygomorphic +  and actinomorphic +
unscented +
1 +  and several +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Sandy soil or dunes in Great Basin Desert with Artemisia, Ericameria, or Sarcobatus. +
suffrutescent +
nodding +  and erect +
13 cm130 mm <br />0.13 m <br /> (26 cm260 mm <br />0.26 m <br />) +
deciduous +
cauline +  and basal +
1 cm10 mm <br />0.01 m <br /> (13 cm130 mm <br />0.13 m <br />) +
deeply sinuate-dentate +  and pinnatifid +
pedicellate +  and sessile +
1 +  and 8 +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
fading pink +  and deep pink +
obcordate +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (8 cm80 mm <br />0.08 m <br />) +
Flowering (Mar–)Jun–Jul(–Sep). +
basal +  and cauline +
clavate +  and globose +
sculptured +  and smooth +
obovoid +
reduced +
2.2 cm22 mm <br />0.022 m <br /> (2.7 cm27 mm <br />0.027 m <br />) +
not spotted +
deciduous +
1.3 cm13 mm <br />0.013 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
with short , lateral branches +  and unbranched +
decumbent +  and ascending or erect +
petiolate +  and sessile +
intrapetiolar +
pubescent +  and glabrous +
3.5 cm35 mm <br />0.035 m <br /> (6 cm60 mm <br />0.06 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
alveolate +
O. trichocalyx var. piperi +
Oenothera deltoides subsp. piperi +
Oenothera deltoides +
subspecies +
3(-5)-aperturate +
0 cm0 mm <br />0 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
woody +  and hard +
smooth +  and erose +
papillate +