Oenothera macrocarpa subsp. fremontii

(S. Watson) W. L. Wagner

Ann. Missouri Bot. Gard. 70: 194. 1983.

Basionym: Oenothera fremontii S. Watson Proc. Amer. Acad. Arts 8: 587. 1873
Synonyms: Megapterium fremontii (S. Watson) Britton
Treatment appears in FNA Volume 10.

Herbs densely strigillose. Stems numerous, with numerous short secondary branches, 3–30 cm. Leaves gray, (2.8–) 3.7–11 × 0.1–0.6 (–1.5) cm; blade linear to narrowly elliptic to narrowly elliptic-lanceolate or narrowly oblanceolate, margins flat, entire or inconspicuously denticulate, apex acute. Flowers: buds with unequal free tips 1–2 (–5) mm; floral-tube (21–) 35–65 (–80) mm; sepals (20–) 25–30 (–37) mm; petals (17–) 25–33 (–37) mm; filaments 13–18 mm, anthers 10–12 mm; style (45–) 55–80 (–98) mm. Capsules ellip­soid to narrowly ellipsoid, often twisted, wings 2–5 (–9) mm wide, body 13–30 (–65) × 2–6 mm. 2n = 14.


Phenology: Flowering May–Aug.
Habitat: Rocky soil derived from fine-textured sandstone, shale or chalk on rocky hillsides, bluffs, badlands.
Elevation: 400–900 m.

Discussion

Subspecies fremontii occurs from Franklin and Webster counties in south-central Nebraska south into Kansas to Ellsworth, Hodgeman, and Logan counties; also with disjunct locations in Antelope and Cedar counties in northeastern Nebraska, and Barber County in south-central Kansas. Some specimens from the eastern part of the range, where subsp. fremontii and subsp. macrocarpa are sympatric, appear intermediate between the two subspecies and are difficult to assign.

Selected References

None.

Lower Taxa

None.

"/4" is not declared as a valid unit of measurement for this property.

2.4 cm24 mm <br />0.024 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Warren L. Wagner +
(S. Watson) W. L. Wagner +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
pedicel-like +
Oenothera fremontii +
indehiscent +  and loculicidal +
lanceolate-elliptic +
pinnatifid +  and dentate +
narrowly elliptic +  and narrowly elliptic-lanceolate or narrowly oblanceolate +
3 cm30 mm <br />0.03 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
quadrangular +
curved +  and straight +
ellipsoid +
Kans. +  and Nebr. +
400–900 m. +
whitish +  and green +
exfoliating +
4 cm40 mm <br />0.04 m <br /> (4.4 cm44 mm <br />0.044 m <br />) +
basifixed +  and versatile +
1.3 cm13 mm <br />0.013 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
6.5 cm65 mm <br />0.065 m <br /> (8 cm80 mm <br />0.08 m <br />) +
hispid +, lanate +  and glabrous +
3.5 cm35 mm <br />0.035 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
zygomorphic +  and actinomorphic +
scented +
2 +  and 1 +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Rocky soil derived from fine-textured sandstone, shale or chalk on rocky hillsides, bluffs, badlands. +
perennial +
suffrutescent +
strigillose +
nodding +  and erect +
2.8 cm28 mm <br />0.028 m <br /> (3.7 cm37 mm <br />0.037 m <br />) +
21 cm210 mm <br />0.21 m <br /> (34 cm340 mm <br />0.34 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
deciduous +
3.7 cm37 mm <br />0.037 m <br /> (11 cm110 mm <br />0.11 m <br />) +
5 cm50 mm <br />0.05 m <br /> (21 cm210 mm <br />0.21 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
denticulate +, entire +  and flat +
pedicellate +  and sessile +
1 +  and 8 +
1.2 cm12 mm <br />0.012 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
3.3 cm33 mm <br />0.033 m <br /> (3.7 cm37 mm <br />0.037 m <br />) +
reddish orange +, fading orange +  and bright-yellow +
obovate +  and obovate to very +
2.5 cm25 mm <br />0.025 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Flowering May–Aug. +
Ann. Missouri Bot. Gard. +
basal +  and cauline +
clavate +  and globose +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
brown +, grayish +  and yellowish-brown brown or dark purplish +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
8 +  and numerous +
sculptured +  and smooth +
obovoid +
reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
3 cm30 mm <br />0.03 m <br /> (3.7 cm37 mm <br />0.037 m <br />) +
purple +, pink +, red +  and green +
deciduous +
coherent +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
40 cm400 mm <br />0.4 m <br /> (60 cm600 mm <br />0.6 m <br />) +
3 cm30 mm <br />0.03 m <br /> (30 cm300 mm <br />0.3 m <br />) +
petiolate +  and sessile +
intrapetiolar +
8 cm80 mm <br />0.08 m <br /> (9.8 cm98 mm <br />0.098 m <br />) +
pubescent +  and glabrous +
5.5 cm55 mm <br />0.055 m <br /> (8 cm80 mm <br />0.08 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
Megapterium fremontii +
Oenothera macrocarpa subsp. fremontii +
Oenothera macrocarpa +
subspecies +
3(-5)-aperturate +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
unequal +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
woody +  and hard +
smooth +  and erose +
not developed +
papillate +
2mm +  and 5mm +