Oenothera macrocarpa subsp. oklahomensis

(Norton) W. L. Wagner

Ann. Missouri Bot. Gard. 70: 194. 1983.

Basionym: Megapteriumoklahomense norton Rep. (Annual) Missouri Bot. Gard. 9: 153, plate 47, figs. 1–3. 1898
Synonyms: Oenothera macrocarpa var. oklahomensis (Norton) Reveal O. missourensis var. oklahomensis (Norton) Munz
Treatment appears in FNA Volume 10.

Herbs glabrous. Stems several, unbranched, sometimes with shorter secondary branches, 5–30 (–42) cm. Leaves green, usually flecked with reddish purple splotches or reddish tinged, (5.5–) 7–11 (–12.5) × (0.3–) 0.8–2 (–3) cm; blade usually elliptic to lanceolate-elliptic, sometimes linear, moderately thick, fleshy, margins usually undu­late, usually conspicuously denticulate to serrulate, rarely subentire, apex acute. Flowers: buds with unequal free tips (5–) 6–10 (–15) mm; floral-tube (82–) 90–140 (–147) mm; sepals 35–60 mm; petals (38–) 42–55 (–65) mm; filaments 24–34 mm, anthers (15–) 19–21 (–25) mm; style (120–) 140–190 mm. Capsules globose to narrowly ellipsoid, not twisted, wings (10–) 15–20 (–25) mm wide, body 35–55 (–75) × 7–9 mm. 2n = 14.


Phenology: Flowering Apr–Jun(–Sep).
Habitat: Rocky, clay soil, open sites of fine-textured, red sandstone usually mixed with gypsum, on pure gypsum, on limestone.
Elevation: 200–500(–800) m.

Discussion

Subspecies oklahomensis is known from Barber, Harper, Meade, and Montgomery counties in extreme southern Kansas south across central Oklahoma to Harmon County in southwestern Oklahoma, and Cooke and Knox counties in adjacent Texas.

Selected References

None.

Lower Taxa

None.

"thick" is not a number."/4" is not declared as a valid unit of measurement for this property.

2.1 cm21 mm <br />0.021 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1.9 cm19 mm <br />0.019 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
Warren L. Wagner +
(Norton) W. L. Wagner +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
pedicel-like +
Megapteriumoklahomense norton +
indehiscent +  and loculicidal +
elliptic to lanceolate-elliptic +  and elliptic +
pinnatifid +  and dentate +
elliptic +  and oblanceolate or suborbiculate +
5.5 cm55 mm <br />0.055 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
3.5 cm35 mm <br />0.035 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
quadrangular +
curved +  and straight +
globose +  and narrowly ellipsoid +
Kans. +, Okla. +  and Tex. +
200–500(–800) m. +
whitish +  and green +
exfoliating +
4 cm40 mm <br />0.04 m <br /> (4.4 cm44 mm <br />0.044 m <br />) +
basifixed +  and versatile +
2.4 cm24 mm <br />0.024 m <br /> (3.4 cm34 mm <br />0.034 m <br />) +
14 cm140 mm <br />0.14 m <br /> (14.7 cm147 mm <br />0.147 m <br />) +
hispid +, lanate +  and glabrous +
9 cm90 mm <br />0.09 m <br /> (14 cm140 mm <br />0.14 m <br />) +
zygomorphic +  and actinomorphic +
scented +
2 +  and 1 +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Rocky, clay soil, open sites of fine-textured, red sandstone usually mixed with gypsum, on pure gypsum, on limestone. +
perennial +
suffrutescent +
nodding +  and erect +
11 cm110 mm <br />0.11 m <br /> (12.5 cm125 mm <br />0.125 m <br />) +
21 cm210 mm <br />0.21 m <br /> (34 cm340 mm <br />0.34 m <br />) +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
reddish tinged +, flecked +  and green +
deciduous +
7 cm70 mm <br />0.07 m <br /> (11 cm110 mm <br />0.11 m <br />) +
5 cm50 mm <br />0.05 m <br /> (21 cm210 mm <br />0.21 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
subentire +, usually conspicuously denticulate +  and serrulate +
pedicellate +  and sessile +
1 +  and 8 +
1.2 cm12 mm <br />0.012 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
5.5 cm55 mm <br />0.055 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
reddish orange +, fading orange +  and bright-yellow +
obovate +  and obovate to very +
4.2 cm42 mm <br />0.042 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
4 cm40 mm <br />0.04 m <br /> (6 cm60 mm <br />0.06 m <br />) +
1 cm10 mm <br />0.01 m <br /> (4 cm40 mm <br />0.04 m <br />) +
Flowering Apr–Jun(–Sep). +
Ann. Missouri Bot. Gard. +
basal +  and cauline +
clavate +  and globose +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
brown +, grayish +  and yellowish-brown brown or dark purplish +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
8 +  and numerous +
sculptured +  and smooth +
obovoid +
reduced +
0.1 cm1 mm <br />0.001 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
6.5 cm65 mm <br />0.065 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
purple +, pink +, red +  and green +
deciduous +
coherent +
3.5 cm35 mm <br />0.035 m <br /> (6 cm60 mm <br />0.06 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
reddish purple +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
30 cm300 mm <br />0.3 m <br /> (42 cm420 mm <br />0.42 m <br />) +
5 cm50 mm <br />0.05 m <br /> (30 cm300 mm <br />0.3 m <br />) +
petiolate +  and sessile +
intrapetiolar +
12 cm120 mm <br />0.12 m <br /> (14 cm140 mm <br />0.14 m <br />) +
pubescent +  and glabrous +
14 cm140 mm <br />0.14 m <br /> (19 cm190 mm <br />0.19 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
Oenothera macrocarpa var. oklahomensis +  and O. missourensis var. oklahomensis +
Oenothera macrocarpa subsp. oklahomensis +
Oenothera macrocarpa +
subspecies +
3(-5)-aperturate +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
unequal +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
woody +  and hard +
smooth +  and erose +
not developed +
papillate +
15mm +  and 20mm +