Oenothera pallida subsp. trichocalyx

(Nuttall) Munz & W. M. Klein in N. L. Britton et al.

N. Amer. Fl., ser. 2, 5: 119. 1965.

Basionym: Oenothera trichocalyx Nuttall in J. Torrey and A. Gray Fl. N. Amer. 1: 494. 1840
Synonyms: Anogra rhizomata A. Nelson A. trichocalyx (Nuttall) Small A. violacea A. Nelson A. vreelandii Rydberg O. albicaulis var. trichocalyx (Nuttall) Engelmann O. pallida var. trichocalyx (Nuttall) Dorn
Treatment appears in FNA Volume 10.
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Herbs usually annual, sometimes perennial, strigillose throughout and villous distally, especially on flower parts; from a taproot, when perennial sometimes lateral roots producing adventitious shoots. Stems single to several from base, usually unbranched. Leaves: basal rosette usually present at anthesis, 3–5 (–7.8) × 0.4–0.8 (–1.2) cm; blade narrowly lanceolate to oblong, margins pinnatifid or dentate. Flowers: buds with free tips 0–0.2 mm; floral-tube 20–30 mm; sepals 10–18 mm; petals 10–20 mm. Capsules spreading to reflexed, straight or contorted. 2n = 14.


Phenology: Flowering (Apr–)May–Jun.
Habitat: Sandy, silty, or rocky soil in pinyon-juniper woodlands or shrublands, with Artemisia and Ericameria.
Elevation: 1100–2500 m.

Distribution

Ariz., Colo., N.Mex., Utah, Wyo.

Discussion

Subspecies trichocalyx occurs across central to southern Wyoming, eastern Utah, western Colorado, northeastern Arizona, and northwestern New Mexico. Within its range it has slight overlap with subspp. pallida and runcinata. In its purest form, subsp. trichocalyx is the most distinctive phase of Oenothera pallida, but many of the populations have character­istics that approach other subspecies with perennial habit (versus annual) appearing occasionally. Plants that are glabrous, or nearly so, like subsp. pallida, but with apparent short-duration habit and divided leaves like subsp. trichocalyx, occur in southern Wyoming and in the Uinta Basin region of Utah; only more typical plants of subsp. trichocalyx otherwise occur in the region without any current evidence of the presence of subsp. pallida.

Selected References

None.

Lower Taxa

None.
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Warren L. Wagner +
(Nuttall) Munz & W. M. Klein in N. L. Britton et al. +
5 cm50 mm <br />0.05 m <br /> (7.8 cm78 mm <br />0.078 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
3 cm30 mm <br />0.03 m <br /> (5 cm50 mm <br />0.05 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
pale green +  and yellow usually fading orange purple pale-yellow reddish or whitish +
pedicel-like +
Oenothera trichocalyx +
indehiscent +  and loculicidal +
sinuate-dentate +  and pinnatifid denticulate subentire or entire +
narrowly lanceolate +  and oblong +
quadrangular +
1.5 cm15 mm <br />0.015 m <br /> (6 cm60 mm <br />0.06 m <br />) +
spreading +  and reflexed +
tapering +, 4-angled +  and cylindrical +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
Ariz. +, Colo. +, N.Mex. +, Utah +  and Wyo. +
1100–2500 m. +
pink +  and white +
exfoliating +
basifixed +  and versatile +
0.9 cm9 mm <br />0.009 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
4 cm40 mm <br />0.04 m <br /> (5 cm50 mm <br />0.05 m <br />) +
hispid +, lanate +  and glabrous +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
zygomorphic +  and actinomorphic +
unscented +
1 +  and several +
curved +  and straight +
indehiscent +  and dehiscent +
4-angled +, terete +, tapering +, cylindrical +, ellipsoid +, clavate +, ovoid ellipsoid +  and cylindrical rhombic-obovoid or globose +
Sandy, silty, or rocky soil in pinyon-juniper woodlands or shrublands, with Artemisia and Ericameria. +
perennial +  and annual +
suffrutescent +
villous +  and strigillose +
nodding +  and erect +
13 cm130 mm <br />0.13 m <br /> (26 cm260 mm <br />0.26 m <br />) +
deciduous +
1 cm10 mm <br />0.01 m <br /> (13 cm130 mm <br />0.13 m <br />) +
dentate +  and pinnatifid +
pedicellate +  and sessile +
1 +  and 8 +
2.5 cm25 mm <br />0.025 m <br /> (4 cm40 mm <br />0.04 m <br />) +
fading pink +  and deep pink +
obcordate +  and obovate +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
2 cm20 mm <br />0.02 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering (Apr–)May–Jun. +
N. Amer. Fl., ser. +
5 cm50 mm <br />0.05 m <br /> (7.8 cm78 mm <br />0.078 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
basal +  and cauline +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
clavate +  and globose +
black +  and brownish +
sculptured +  and smooth +
obovoid +
reduced +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
not spotted +
deciduous +
1 cm10 mm <br />0.01 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
parietal +, axile +  and placentation +
unequal +
adventitious +
aquatic +, amphibious +  and terrestrial +
in 2 unequal series +  and subequal +
2 times as many or as many as sepals +
50 cm500 mm <br />0.5 m <br /> (70 cm700 mm <br />0.7 m <br />) +
ascending +  and erect +
single +  and several +
10 cm100 mm <br />0.1 m <br /> (50 cm500 mm <br />0.5 m <br />) +
petiolate +  and sessile +
intrapetiolar +
pubescent +  and glabrous +
2.5 cm25 mm <br />0.025 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
aquatic +, amphibious +  and terrestrial +
smooth +, reticulate +  and furrowed +
alveolate +
Anogra rhizomata +, A. trichocalyx +, A. violacea +, A. vreelandii +, O. albicaulis var. trichocalyx +  and O. pallida var. trichocalyx +
Oenothera pallida subsp. trichocalyx +
Oenothera pallida +
subspecies +
3(-5)-aperturate +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
woody +  and hard +
smooth +  and erose +
papillate +