Orobanche californica subsp. grandis

Heckard

Madroño 22: 60, figs. 1P–R, 3A, 4E. 1973.

Endemic
Synonyms: Aphyllon californicum subsp. grande (Heckard) A. C. Schneider
Treatment appears in FNA Volume 17. Treatment on page 481. Mentioned on page 478, 479.

Plants simple, rarely branched from base, 8–35 cm, proximal portion to inflorescence (5–) 10–20 cm, stout. Inflorescences corymbs, sometimes subcapitate to subcorymbose racemes, 3–15 cm; bracts pallid to pinkish tinged, drying brown. Pedicels 5–15 (–25) mm. Calyces 12–20 (–25) mm, lobes buff or yellow, tinged pink or brownish, linear-subulate, 15–20 mm. Corollas pallid to pinkish tinged, drying reddish-brown, 35–50 mm; tube broad, abruptly widening toward throat; throat 9–10 mm wide; lips 12–14 mm, abaxial lobes lanceolate to lanceolate-ovate, 5–7 mm wide, apex acute, adaxial lobes ovate, apex obtuse, shallowly notched, erose.


Phenology: Flowering Apr–Oct(–Nov).
Habitat: Sandy soils or lee of foredunes near coast.
Elevation: 0–150 m.

Discussion

Subspecies grandis is commonly found near the ocean shore from Los Angeles to San Mateo counties and on Santa Rosa Island. It is uncommon, and its habitat is impacted by Carpobrotus (Aizoaceae) invasions. The primary host is Corethrogyne filaginifolia (Asteraceae), but it can also utilize Grindelia (Asteraceae). Other hosts reported are Adenostoma fasciculatum (Rosaceae), Artemisia, Isocoma veneta, and Heterotheca (Asteraceae).

Plants included in subsp. grandis have been identified variously in the past. The northern coastal specimens were aligned with subsp. californica, whereas the southern collections were identified as subspp. feudgei or jepsonii. Subspecies grandis is roughly sympatric with other coastal subspecies, but its unique morphology and its preferences for sand dune habitat and Corethrogyne filaginifolia as a host make it distinctive.

Selected References

None.

Lower Taxa

None.

lanceolate +  and lanceolate-ovate +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
glabrous +  and villous +
glandular-pubescent +
Notched (?) +, Obtuse (?) +  and Acute (?) +
L. Turner Collins +, Alison E. L. Colwell +  and George Yatskievych +
Heckard +
enlarged +
oblong;lanceolate;oblong;lanceolate;deltate;triangular;ovate +
0.4 cm4 mm <br />0.004 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
not leathery +, not +  and fleshy +
brown;pallid;pinkish tinged +
appressed +  and spreading +
less lanceolate +  and oblanceolate +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
pallid +  and dark purple pink yellow or white +
divided +
1.2 cm12 mm <br />0.012 m <br /> (2 cm20 mm <br />0.02 m <br />) +
ovoid;cylindric-ovoid +
1 cm10 mm <br />0.01 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
5 cm50 mm <br />0.05 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
reddish-brown +, pallid +  and pinkish tinged +
Bent (?) +  and Tubular (?) +
constricted +
3.5 cm35 mm <br />0.035 m <br /> (5 cm50 mm <br />0.05 m <br />) +
spikelike +
0–150 m. +
pubescent +  and glabrous +
indehiscent +, septicidal +, loculicidal +  and dehiscence +
Sandy soils or lee of foredunes near coast. +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
axillary +  and terminal +
10 cm100 mm <br />0.1 m <br /> (20 cm200 mm <br />0.2 m <br />) +
branched +  and simple +
pallid cream +  and nearly white +
3 cm30 mm <br />0.03 m <br /> (15 cm150 mm <br />0.15 m <br />) +
deciduous +
cream +  and pinkish or purplish tinged or pink +
1.2 cm12 mm <br />0.012 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
brownish +, tinged pink +, yellow +  and buff +
loosely ascending +  and recurved +
glandular-pubescent +
linear-subulate +
erosulate +  and entire +
lobed +, toothed +  and entire +
tenuinucellate +  and unitegmic +
campylotropous-like +
spikelike +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Flowering Apr–Oct(–Nov). +
subcorymbose +
spikelike +
branched +  and unbranched +
inconspicuous +
coralloid +
stout +  and slender +
500 +  and 2000 +
prismatic +, ovoid +  and oblong-ellipsoid +
5 +  and 4 +
subequal +
yellow +  and white +
aerial +  and subterranean +
capitate +  and crateriform-peltate +
clavate +  and 2-4-lobed +
Aphyllon californicum subsp. grande +
Orobanche californica subsp. grandis +
Orobanche californica +
subspecies +
9mm;10mm +
cream;pinkish or purplish tinged or pink +
glandular-puberulent +
constricted +
widening +  and broad +
branched +  and simple +
perennial +  and annual +
holoparasitic +  and hemiparasitic +
stout +  and slender +